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false candytuft

pearshape smelowskia

Habit Plants sometimes canescent basally; caudex usually simple, rarely branched. Plants sometimes canescent throughout; caudex simple.
Stems

several from base, often unbranched or branched proximally, 0.6–3(–3.5) dm, trichomes simple, to 1 mm, mixed with smaller, dendritic ones.

usually several, rarely simple from base, usually unbranched, rarely branched, 0.5–2.4(–3) dm, trichomes simple, to 1.3 mm, mixed with smaller, dendritic ones.

Basal leaves

petiole 0.5–3.2 cm, ciliate or not;

blade obovate to ovate in outline, (terminal segments linear, oblong, or ovate), 0.7–2.5 cm × 4–14 mm, (terminal segments 0.2–1.1 cm × 1–5 mm), margins palmately (3 or) 5 (or 7)-lobed, (surfaces densely pubescent, trichomes dendritic, often mixed with larger, simple ones, to 1 mm), apex obtuse or subacute.

petiole 1–4 cm, often ciliate, trichomes simple;

blade broadly ovate to oblong in outline, (terminal segments oblong to or ovate), 1–2.2 cm × 8–15 mm, (terminal segments 0.4–1 cm × 1.5–3 mm), margins pinnatifid, apex obtuse.

Cauline leaves

shortly petiolate or sessile;

blade often similar to basal, smaller distally, margins sometimes pinnatifid.

shortly petiolate or sessile;

blade similar to basal, smaller distally.

Racemes

considerably elongated in fruit.

considerably elongated in fruit.

Flowers

sepals (usually persistent), 2.5–3.2 mm;

petals purple to lavender, suborbicular to obovate, 4–5 × 1.5–2 mm, narrowed to claw, 1.5–2.5 mm, apex rounded;

anthers oblong, 0.4–0.5 mm.

sepals 1.2–2.2 mm;

petals purple, lavender, or white, suborbicular to obovate, 2.5–4.5 × 1–2.5 mm, narrowed to claw, 1–2 mm, apex rounded;

anthers ovate, 0.3–0.5 mm.

Fruiting pedicels

spreading to divaricate-ascending, (secund, often curved), proximalmost bracteate, 6–15(–20) mm, pubescent, trichomes simple mixed with smaller, dendritic ones.

ascending to divaricate-ascending, (often forming less than 40˚ angle), proximalmost bracteate, 5–15(–20) mm, pubescent, trichomes simple (to 1.4 mm), mixed with smaller, dendritic ones.

Fruits

spreading to ascending, usually oblanceolate to spatulate or linear-oblanceolate, rarely oblong-obovate, angustiseptate, (8–)12–28 × 4–8 mm, base cuneate, apex obtuse;

valves each with prominent midvein and distinct lateral veins, (rarely sparsely pubescent);

ovules 10–18 per ovary;

style 0.1–1.1 mm.

ascending, pyriform, subterete, 5–9 × 2.5–4 mm, base cuneate, apex rounded;

valves each with prominent midvein;

ovules 4 per ovary;

style 0.4–1.2 mm.

Seeds

1.5–2.2 × 1–1.3 mm.

(oblong-lanceolate) 2.5–3.2 × 1–1.5 mm.

2n

= 12.

= 12.

Smelowskia borealis

Smelowskia pyriformis

Phenology Flowering Jun–Aug. Flowering Jun–Aug.
Habitat Loose talus, metamorphic slide rock, rocky slopes, scree, shale splinters, limestone rubble, alpine ridges, barren rocks, unstable talus in alpine glacial bowl Loose talus, scree slopes, shale ridgetops, limestone volcanic rubble, mixed sandstone-siltstone-carbonate scree
Elevation 600-1700 m (2000-5600 ft) 600-1700 m (2000-5600 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
from FNA
AK
[BONAP county map]
Discussion

W. H. Drury Jr. and R. C. Rollins (1952) and Rollins (1993) divided Smelowskia borealis into four varieties based on the duration of sepals (persistent versus caducous), style length (less than versus greater than 0.5 mm), and density of indumentum. The various states of these three characters are present in all possible combinations, and they are not inherited in the combinations presented by these authors. Some forms of the species are densely villous and resemble S. johnsonii in their leaf indumentum and perhaps merit recognition as an infraspecific taxon (var. villosa). As delimited here, S. borealis is quite variable; more studies are needed to determine whether or not any of the four varieties merit recognition at some rank. Although numerous collections of the species were examined, none of the types was available for preparing this account. The species is easily distinguished from the remaining North American Smelowskia by having secund infructescences and distinctly angustiseptate fruits.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 673. FNA vol. 7, p. 675.
Parent taxa Brassicaceae > tribe Smelowskieae > Smelowskia Brassicaceae > tribe Smelowskieae > Smelowskia
Sibling taxa
S. americana, S. johnsonii, S. media, S. ovalis, S. porsildii, S. pyriformis
S. americana, S. borealis, S. johnsonii, S. media, S. ovalis, S. porsildii
Synonyms Melanidion boreale, Acroschizocarpus kolianus, Ermania borealis, S. borealis var. jordalii, S. borealis var. koliana, S. borealis var. villosa
Name authority (Greene) W. H. Drury & Rollins: Rhodora 54: 111. (1952) W. H. Drury & Rollins: Rhodora 54: 108, fig. 3B. (1952)
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