Sisymbrium officinale |
Sisymbrium |
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common hedge-mustard, common tumble-mustard, hedge mustard |
hedge-mustard, rocket, tumblemustard |
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Habit | Annuals; glabrous or pubescent. | Plants not scapose; pubescent or glabrous. | ||||||||||||||||||||||||||||
Stems | erect, branched distally, 2.5–7.5(–11) dm, usually sparsely to densely hirsute, (trichomes retrorse), rarely glabrate distally. |
often erect, sometimes ascending, rarely subprostrate or decumbent, often branched distally, sometimes unbranched. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins dentate, sinuate, lyrate, runcinate, or pinnately lobed [entire]; cauline similar to basal, (blade smaller distally). |
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Basal leaves | usually rosulate; petiole (1–)2–7(–10) cm; blade broadly oblanceolate or oblong-obovate (in outline), (2–)3–10(–15) cm × (10–)20–50(–80) mm, margins lyrate-pinnatifid, pinnatisect, or runcinate; lobes (2)3 or 4(5) on each side, oblong or lanceolate, smaller than terminal lobe, margins entire, dentate, or lobed, (terminal lobe suborbicular or deltate, margins dentate). |
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Cauline leaves | similar to basal; blade with lobe margins dentate or subentire. |
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Racemes | (several-flowered), often considerably elongated in fruit. |
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Flowers | sepals erect, oblong-ovate, 2–2.5 × ca. 1 mm; petals spatulate, 2.5–4 × 1–2 mm, claw 1–2 mm; filaments (erect, yellowish), 2–3 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate or oblong, (glabrous or pubescent); petals yellow, obovate, spatulate, oblong, or suborbicular, (longer than sepals), claw differentiated from blade, (subequaling or longer than sepals, apex obtuse or emarginate); stamens tetradynamous; filaments not dilated basally; anthers oblong, (apex obtuse); nectar glands confluent, subtending bases of stamens, median glands present. |
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Fruiting pedicels | erect, (appressed to rachis), stout, narrower than fruit, 1.5–3(–4) mm. |
ascending, divaricate, or erect, slender or stout (sometimes as wide as fruit). |
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Fruits | (erect), subulate-linear, straight, slightly torulose or smooth, stout, (0.7–)1–1.4(–1.8) cm × 1–1.5 mm; valves glabrous or pubescent; ovules 10–20 per ovary; style (0.8–)1–1.5(–2) mm; stigma slightly 2-lobed. |
usually sessile, rarely shortly stipitate (gynophore to 1 mm), usually linear, rarely lanceolate or subulate, smooth or torulose; valves each with prominent midvein and 2 conspicuous marginal veins, usually glabrous, rarely pubescent; replum rounded; septum complete; style subclavate [clavate, conical, cylindrical]; stigma capitate (lobes not decurrent). |
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Seeds | 1–1.3 × 0.5–0.6 mm. |
plump, not winged, oblong [ovoid]; seed coat (reticulate or papillate), not mucilaginous when wetted; cotyledons incumbent. |
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x | = 7. |
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2n | = 14. |
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Sisymbrium officinale |
Sisymbrium |
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Phenology | Flowering Apr-late Sep. | |||||||||||||||||||||||||||||
Habitat | Roadsides, fields, pastures, waste grounds, deserts | |||||||||||||||||||||||||||||
Elevation | 0-2200 m (0-7200 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; CA; CT; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; YT; Europe; Asia; n Africa [Introduced in North America; introduced also in Central America, South America, Australia]
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North America; Europe; Asia; Africa [Introduced in Central America, South America, Australia] |
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Discussion | Species 41 (8 in the flora). All except one of the eight species of Sisymbrium in North America are introduced Eurasian weeds. Sisymbrium linifolium (Nuttall) Nuttall was retained in Sisymbrium by both E. B. Payson (1922) and O. E. Schulz (1924). Greene took that species as the type of his genus Schoenocrambe. R. C. Rollins (1982b, 1993) maintained Schoenocrambe and (1993) recognized Sisymbrium auriculatum as the only native North American species of the genus. Molecular studies (S. I. Warwick et al. 2002, 2005) clearly demonstrated that Schoenocrambe should be united with Sisymbrium, that S. linifolium is most closely related to the Eurasian S. polymorphum (Murray) Roth (as was suggested by both Payson and Schulz), that 40 of the 41 species of Sisymbrium are native to the Old World, and that S. auriculatum is a member of the New World Thelypodieae and is unrelated to Sisymbrium. See also Warwick and I. A. Al-Shehbaz (2003) and Al-Shehbaz (2005). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 670. | FNA vol. 7, p. 667. | ||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Sisymbrieae > Sisymbrium | Brassicaceae > tribe Sisymbrieae | ||||||||||||||||||||||||||||
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Synonyms | Erysimum officinale, S. officinale var. leiocarpum | Chamaeplium, Norta, Pachypodium, Schoenocrambe, Velarum | ||||||||||||||||||||||||||||
Name authority | (Linnaeus) Scopoli: Fl. Carniol. ed. 2, 2: 26. (1772) | Linnaeus: Sp. Pl. 2: 657. (1753) | ||||||||||||||||||||||||||||
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