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Dolores campion, San Francisco campion

dusty-miller, lychnide coronaire, mullein pink, rose campion

Habit Plants perennial; taproot stout; caudex branched, woody. Plants perennial, grayish white-tomentose, eglandular; taproot slender to stout; caudex branched, slightly woody.
Stems

usually several–many, rarely 1, erect, leafy, 10–55 cm, base often decumbent with marcescent leaf bases, scabrous-puberulent to pubescent, usually ± viscid-glandular distally, rarely densely so.

several, erect, branched distally, stout, 40–100 cm.

Leaves

2 per node;

basal petiolate, blade linear-lanceolate to lanceolate, 3–10 cm × 2–13 mm (including petiole), apex acute, glabrous to scabrous-puberulent or softly pubescent, petiole often ciliate;

cauline sessile or petiolate, connate proximally, reduced distally, blade linear to narrowly lanceolate, 2–10 cm × 2–8 mm.

basal blade oblanceolate, spatulate, 5–10 cm × 10–25(–30) mm, margins entire, apex acute, apiculate, with tuft of white hairs;

cauline in 5–10 pairs, sessile, reduced distally, blade with both surfaces obscured by dense, silky, grayish-white tomentum.

Inflorescences

cymose with elongate ascending branches, open, flowers (1–)3 to many, bracteate, bracteolate;

bracts and bracteoles lanceolate, 3–20 mm, apex acuminate.

with 1–several dichotomies, several-flowered, open, bracteate;

branches ascending, elongate;

bracts leaflike, 10–20 mm.

Pedicels

1/4–3 times longer than calyx, scabrous-puberulous to pubescent and glandular, ± viscid.

straight, stout, to 10 cm.

Flowers

calyx prominently 10-veined, tubular in flower, ± clavate and contracted around carpophore in fruit, 10–14 × 4–6 mm, margins dentate, veins parallel, green (rarely purplish), with pale commissures, lobes ascending, lanceolate, 2–3 mm, margins usually with obtuse, membranous border, apex spreading, shortly glandular-pubescent, usually viscid;

corolla off-white (greenish) to dusky pink, clawed, claw equaling calyx, limb obovate, 2-lobed, 3–7 mm, shorter than claw, rarely with small lateral teeth, appendages 2, usually lacerate, 1–2 mm;

stamens ca. equaling petals;

styles 3(–4), often much longer than petals.

ca. 35 mm;

calyx thickly 10-veined, obovate, ca. 15 × 10 mm in fruit, margins dentate with 5 narrowly lanceolate lobes ca. 1/4 length of tube, tomentose;

corolla rich magenta-pink, sometimes white, clawed, claw equaling calyx, limb spreading horizontally, broadly obovate, shallowly 2-lobed, appendages 2, narrow, 2–4 mm;

stamens equaling claw;

stigmas 5, equaling claw.

Capsules

ovoid-ellipsoid, slightly longer than calyx, opening by 6 (or 8) recurved teeth;

carpophore 2–5 mm.

equaling to tightly enclosed within calyx, obovate-elliptic, ca. 14 mm, opening by 5 spreading, lanceolate teeth;

carpophore ca. 2 mm.

Seeds

dark reddish brown to black, reniform, ca. 1.5 mm, papillate, with larger papillae around margins.

grayish brown, reniform-rotund, plump, 1–1.5 mm, coarsely verrucate.

2n

= 48.

= 24.

Silene verecunda

Silene coronaria

Phenology Flowering summer. Flowering summer.
Habitat Meadows, chaparral, sagebush, open woodlands, dry pine forests, alpine ridges, dry canyons Roadsides, fields, waste or rocky places
Elevation 0-3400 m (0-11200 ft) 0-300 m (0-1000 ft)
Distribution
from FNA
AZ; CA; NV; OR; UT; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; ID; IL; IN; LA; MA; MD; MI; MO; NC; NH; NJ; NY; OH; OR; PA; SC; UT; VA; VT; WA; WI; WV; BC; NB; NS; ON; QC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene verecunda is an exceptionally variable species, very difficult to circumscribe and tending to intergrade with S. bernardina, S. oregana, and S. grayi. It differs from the first two of those species mainly in having two-lobed petals. Silene grayi is a much smaller cespitose alpine plant with very large seeds. Hitchcock and Maguire divided S. verecunda into subsp. verecunda, subsp. platyota, and subsp. andersonii. Of these, subsp. andersonii is the most distinct, with a scabrous-puberulent indumentum, very narrow, stiff leaves, and rigid stems that are decumbent at the base, with marcescent leaf bases. The claw of the petals also is often more uniformly ciliate. Subspecies verecunda has a very different appearance, its mature calyx being shorter, broader, and markedly clavate. It is a short, stocky, viscid-glandular plant of exposed coastal habitats and may simply be a local ecotype. Subspecies platyota encompasses the remainder of the variation in the complex. Most of this variation consists of plants with fewer flowering stems; softer pubescence; broader, flat leaves; and thinner, more papery calyces. All these forms of S. verecunda appear to intergrade freely and, based on current information, any separation would be arbitrary. The species is in need of an experimental study to determine the nature of variation and its taxonomic value.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene coronaria is commonly cultivated and occasionally escapes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 211. FNA vol. 5, p. 180.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Synonyms S. andersonii, S. behrii, S. luisana, S. occidentalis var. nancta, S. platyota, S. verecunda subsp. andersonii, S. verecunda var. eglandulosa, S. verecunda subsp. platyota, S. verecunda var. platyota Agrostemma coronaria, Lychnis coronaria
Name authority S. Watson: Proc. Amer. Acad. Arts 10: 344. (1875) (Linnaeus) Clairville: Man. Herbor. Suisse, 145. (1811)
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