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apetalous catchfly, mountain campion, nodding campion, Spalding's campion

Parish's campion, Parish's catchfly

Habit Plants perennial, cespitose or not; taproot stout. Plants perennial; taproot stout; caudex much-branched, woody.
Stems

erect, branched or simple, 5–40 cm, glabrous or pubescent.

many, decumbent to erect, usually simple, 10–40 cm, woody, pilose and glandular (rarely eglandular proximally).

Leaves

basal few or numerous, petiolate, blade narrowly oblanceolate, spatulate, 1–13 cm × 1–7 mm, margins ciliate, glabrous or pubescent;

cauline in 1–5 pairs, sessile, much reduced distally, blade linear to lanceolate, 0.1–4 cm × 1–5 mm.

in 5–8 pairs, sessile but proximal ones narrowed into pseudopetiole, largest in mid-stem region, reduced proximally, scalelike at base, blade narrowly lanceolate to oblanceolate-elliptic or obovate, 2–6 cm × 3–20 mm, apex acute and acuminate, usually thick, leathery, densely puberulent and viscid-glandular or eglandular.

Inflorescences

cymose, open or congested, 3–15(–30)-flowered, sometimes compound, leafy.

Pedicels

slender or stout.

to 1(–1.5) cm, shortly pilose, viscid-glandular, flowers sometimes sessile.

Flowers

erect or nodding;

calyx prominently 10-veined, ovate-elliptic to broadly campanulate, inflated, 11–18 × 6–13 mm, thin and papery, veins purple or brown, lobes ovate to triangular, 2–3 mm;

corolla dingy pink, purple, or red, claw shorter than to equaling calyx, limb not differentiated from claw, ovate or obovate, unlobed to 2-lobed, 1–4 mm;

stamens shorter than petals;

styles 5, shorter than petals.

calyx prominently 10-veined, tubular, clavate in fruit, constricted proximally around carpophore, (20–)25–30 × 4–7 mm, papery, densely glandular-puberulent, viscid, veins parallel, green, with pale commissures, lobes narrowly lanceolate, acuminate, 5–8 mm, herbaceous;

corolla pale greenish yellow to white, clawed, claw equaling calyx, ligulate, as broad as limb, limb 7–8 mm, deeply laciniate into 6 or more linear lobes, appendages oblong, laciniate, 2 mm;

stamens equaling calyx;

styles 3, exserted.

Capsules

equaling to slightly longer than calyx, opening by 10 recurved teeth;

carpophore 1–2 mm.

equaling calyx, opening by 6 ascending teeth;

carpophore ca. 3 mm.

Seeds

brown or sooty brown, broadly winged, round to ± angular, 1.5–2.5 mm diam. including wing.

brown, reniform, 1.5–2 mm, margins papillate;

papillae large, inflated.

2n

= 48.

Silene uralensis

Silene parishii

Phenology Flowering spring and summer.
Habitat Rocky ledges and slopes, stream banks, open coniferous woodlands
Elevation 1400-3400 m (4600-11200 ft)
Distribution
from FNA
AK; CO; MT; UT; WY; AB; BC; MB; NL; NT; NU; ON; QC; YT; Canada; Greenland; nw United States including Alaska; e Asia (Russian Far East, Siberia)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Silene uralensis is a very variable species complex. Recently, A. Kurtto (2001) has resurrected S. wahlbergella as a species distinct from S. uralensis, differing in its shorter petals and larger capsule. It appears to be confined to Scandanavia and adjoining arctic Russia. North American reports of S. wahlbergella (and its synonym Lychnis apetala) are all referable to S. uralensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene parishii varies considerably in pubescence and leaf shape. C. L. Hitchcock and B. Maguire (1947) recognized three varieties on the basis of this variation. However, the characters vary independently and have only a weak geographical correlation. Hence recognition of the three varieties serves little useful purpose.

Silene parishii is confined to the mountains of southern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences usually simple, pedicels slender with single nodding (deflexed) flower (fruiting pedicels erect); seeds 1.5-2(-2.5) mm diam. (incl. broad wing).
subsp. uralensis
1. At least some of the inflorescences branched with 2 to several flowers, pedicels erect to angled but not deflexed except at tip in flower; seeds 2-2.5 mm diam
→ 2
2. Stems usually glabrous to sparsely, rarely pubescent, (15-)20-40 cm, slender; inflorescences branched with (1-)2-10 flowers; corolla only slightly exceeding calyx
subsp. ogilviensis
2. Stems densely pubescent with purple-septate hairs, 10-35 cm, stout; at least some inflorescences forked with 1-3(-4) flowers; corolla usually ca. 11/ 2 times calyx
subsp. porsildii
Source FNA vol. 5, p. 209. FNA vol. 5, p. 197.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. uralensis subsp. ogilviensis, S. uralensis subsp. porsildii, S. uralensis subsp. uralensis
Synonyms Gastrolychnis uralensis S. parishii var. latifolia, S. parishii var. viscida
Name authority (Ruprecht) Bocquet: Candollea 22: 25. (1967) S. Watson: Proc. Amer. Acad. Arts 17: 366. (1882)
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