Silene suksdorfii |
Silene bernardina |
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Cascade alpine campion, Cascade alpine catchfly, Cascade catchfly, Suksdorf's catchfly, Suksdorf's silene |
mountain catchfly, Palmer's campion, Palmer's catchfly |
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Habit | Plants perennial, cespitose, with decumbent subterranean shoots; taproot stout; caudex branched, woody. | Plants perennial, loosely cespitose; taproot stout; caudex branched, woody, bearing tufts of leaves. |
Stems | numerous, erect, simple, 3–15 cm, pubescent, viscid-glandular distally. |
not much- branched, slender, (15–)30–60 cm, sparsely pubescent proximally, viscid-glandular distally. |
Leaves | mostly basal, densely tufted; basal numerous, pseudopetiolate, blade narrowly oblanceolate, tapering into base, 0.5–3 cm × 1.5–4 mm, ± fleshy, apex acute, puberulent; cauline in 1–3 pairs, ± sessile, reduced, blade narrowly oblanceolate to linear-lanceolate, 0.7–2 cm × 1–3 mm, apex acute, puberulent. |
mostly basal; blade linear-lanceolate to oblanceolate, 2–8 cm × 2–6(–15) mm (including petiole), base tapered into slender petiole, apex acute to obtuse, subglabrous to glandular-pubescent on both surfaces; cauline leaves to 4 pairs below inflorescence, narrower than basal leaves, blade usually linear but rarely elliptic-lanceolate. |
Inflorescences | flowers terminal, solitary, or in single dichotomy, bracteate; bracts leaflike, 3–15 mm. |
erect, with several short, ascending branches, few-flowered, open, bracteate, shortly pubescent and viscid-glandular; bracts narrowly lanceolate, 3–10 mm, rigid. |
Pedicels | erect, ca. equaling calyx, viscid glandular-pubescent, hairs with purple septa. |
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Flowers | calyx prominently 10-veined, campanulate, not contracted proximally around carpophore, 10–15 × 5–7 mm, papery, veins parallel, purplish, with pale commissures, with purple-septate glandular hairs (rarely septa not purple), lobes ovate, ca. 2 mm, margins broad, membranous, apex obtuse; corolla off-white or tinged with dusky purple, clawed, claw equaling calyx, broadened distally, limb 2-lobed, 3–5 mm, appendages ca. 1 mm; stamens equaling calyx; styles 3(–4), equaling calyx. |
calyx prominently 10-veined, broadly tubular, umbilicate, moderately or not clavate, narrowed around carpophore, lobed, 12–15 × 4–6 mm, thin and papery, with short glandular-viscid pubescence, veins parallel, usually red pigmented, with pale commissures; lobes lanceolate, 2–4 mm, apex acute; petals white, pink, or dingy red, 11/2–2 times calyx, claw equaling calyx, ciliate at base, limb obtriangular, 4–6 mm, deeply divided into 4 linear lobes, appendages 2, conspicuous, laciniate, 2–3 mm, apex rounded; stamens slightly exserted; filaments ciliate at base; styles 3(–4), equaling or longer than stamens. |
Capsules | equaling calyx, opening by 6 (or 8) teeth; carpophore 2.5–3.5 mm. |
1-locular, narrowly ovoid, exceeding calyx, opening by 6 (or 8) ascending teeth; carpophore 3–6 mm. |
Seeds | brown, broadly winged, reniform, 1–2 mm, rugose-tessellate. |
brown, reniform, 1.5–2 mm broad, shallowly tuberculate on both surfaces, papillate around margins. |
2n | = 48. |
= 48. |
Silene suksdorfii |
Silene bernardina |
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Phenology | Flowering summer. | Flowering summer. |
Habitat | Alpine ridges, gravel slopes, talus | Dry, grassy or gravelly slopes, open woodlands |
Elevation | 1600-3000 m (5200-9800 ft) | 1300-3600 m (4300-11800 ft) |
Distribution |
CA; OR; WA
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CA; ID; NV; OR; WA; Mexico (Baja California)
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Discussion | Silene suksdorfii appears to be closely related to S. parryi but differs in its broadly winged seeds, smaller size, cespitose habit, and the prominent purple-septate hairs of the calyx, although the latter occasionally are present in S. parryi. It is very similar to, and in Idaho appears to intergrade with, another alpine species, S. sargentii, which has linear leaves and lacks the purple septa in the hairs and the broad wing on the seeds. It is similar also to S. hitchguirei; see discussion under that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Silene bernardina is the earliest valid name for this species. Watson had previously (1875) named it S. montana, and that name was taken up by C. L. Hitchcock and B. Maguire (1947), who cited S. bernardina as a subspecies of S. montana. Unfortunately, the epithet montana is pre-occupied in Silene by S. montana Arrondeau (1863), an unrelated European species. The situation was further complicated by Watson in 1877, when he used the name Lychnis montana for another unrelated species now transferred to Silene and called S. hitchguirei. Silene bernardina varies in leaf width, pubescence, and flower color. The broader-leaved and more sparsely pubescent forms have been referred to subsp. bernardina, and the more-common, narrower-leaved, more-densely pubescent, and viscid forms have been referred to subsp. maguirei. Some forms of Silene bernardina can be difficult to distinguish from S. verecunda, S. sargentii, and S. oregana. Silene verecunda differs in its smaller, clavate calyx and in its petals being only shortly two-lobed. Silene sargentii is a small, densely cespitose, high-alpine species with very narrow, linear leaves (1–2 mm wide), shortly two-lobed petals, and seeds with much larger papillae around the margins. In S. oregana the petals are larger (two times the calyx) and deeply divided into many very narrow segments; the claw and the filaments are glabrous; the leaves, particularly the basal ones, are broader; and the inflorescences are narrower, with the more numerous flowers arranged on short, ascending branches; also, the calyx lobes are ovate and obtuse instead of lanceolate and acute. The Idaho material tends to be intermediate with S. oregana but has open, dichotomously branched inflorescences, and the petals are nearer to those of S. bernardina. These plants from Valley County in the Payette National Forest need further study, preferably in the field. They may represent a distinct taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 209. | FNA vol. 5, p. 175. |
Parent taxa | Caryophyllaceae > subfam. Caryophylloideae > Silene | Caryophyllaceae > subfam. Caryophylloideae > Silene |
Sibling taxa | ||
Synonyms | S. bernardina subsp. maguirei, S. bernardina var. rigidula, S. bernardina var. sierrae, S. occidentalis var. nancta, S. shockleyi | |
Name authority | B. L. Robinson: Bot. Gaz. 16: 44, plate 6, figs. 9–11. (1891) | S. Watson: Proc. Amer. Acad. Arts 24: 82. (1889) |
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