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Cascade alpine campion, Cascade alpine catchfly, Cascade catchfly, Suksdorf's catchfly, Suksdorf's silene

Habit Plants perennial, cespitose, with decumbent subterranean shoots; taproot stout; caudex branched, woody. Herbs, annual, biennial, or perennial; taprooted or rhizomatous, sometimes stoloniferous.
Stems

numerous, erect, simple, 3–15 cm, pubescent, viscid-glandular distally.

erect or ascending, seldom sprawling, decumbent, or prostrate, simple or branched.

Leaves

mostly basal, densely tufted;

basal numerous, pseudopetiolate, blade narrowly oblanceolate, tapering into base, 0.5–3 cm × 1.5–4 mm, ± fleshy, apex acute, puberulent;

cauline in 1–3 pairs, ± sessile, reduced, blade narrowly oblanceolate to linear-lanceolate, 0.7–2 cm × 1–3 mm, apex acute, puberulent.

opposite, rarely whorled, connate proximally, petiolate (basal leaves) or often sessile, not stipulate;

blade linear or subulate to ovate, not succulent or rarely so (Silene).

Inflorescences

flowers terminal, solitary, or in single dichotomy, bracteate;

bracts leaflike, 3–15 mm.

terminal cymes, thyrses, fascicles, or capitula, or flowers solitary, axillary;

bracts foliaceous, scarious, or absent;

involucel bracteoles present or often absent.

Pedicels

erect, ca. equaling calyx, viscid glandular-pubescent, hairs with purple septa.

present or rarely flowers sessile or subsessile.

Flowers

calyx prominently 10-veined, campanulate, not contracted proximally around carpophore, 10–15 × 5–7 mm, papery, veins parallel, purplish, with pale commissures, with purple-septate glandular hairs (rarely septa not purple), lobes ovate, ca. 2 mm, margins broad, membranous, apex obtuse;

corolla off-white or tinged with dusky purple, clawed, claw equaling calyx, broadened distally, limb 2-lobed, 3–5 mm, appendages ca. 1 mm;

stamens equaling calyx;

styles 3(–4), equaling calyx.

bisexual or seldom unisexual (the species then often dioecious), often conspicuous;

perianth and androecium hypogynous;

sepals 5, connate (1/4–)1/2+ their lengths into cup or tube, (1–)5–40(–62) mm, apex not hooded or awned;

petals absent or 5, often showy, white to pink or red, usually clawed, auricles absent or sometimes present, coronal appendages sometimes present, blade apex entire or emarginate to 2-fid, sometimes dentate to lacinate;

stamens (5 or) 10 (absent in pistillate flowers), in 1 or 2 whorls, arising from base of ovary;

staminodes absent or rarely 1–10;

ovary 1-locular, sometimes 2-locular proximally (Vaccaria), or 3–5-locular (some Silene);

styles 2–3(–5) (absent in staminate flowers), distinct;

stigmas 2–3(–5) (absent in staminate flowers).

Fruits

capsules, opening by 4–6(–10) valves or teeth;

carpophore usually present.

Capsules

equaling calyx, opening by 6 (or 8) teeth;

carpophore 2.5–3.5 mm.

Seeds

brown, broadly winged, reniform, 1–2 mm, rugose-tessellate.

4–150(–500+), reddish to gray or often brown or black, usually reniform and laterally compressed to globose, sometimes oblong or shield-shaped and dorsiventrally compressed;

embryo peripheral and curved, or central and straight.

x

= 7, 10, 12, [13?,] 14, 15, 17, [18].

2n

= 48.

Silene suksdorfii

Caryophyllaceae subfam. caryophylloideae

Phenology Flowering summer.
Habitat Alpine ridges, gravel slopes, talus
Elevation 1600-3000 m (5200-9800 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
North-temperate regions; Europe (esp Mediterranean region); Asia (esp Mediterranean region e to c Asia); Africa (Mediterranean region, Republic of South Africa)
Discussion

Silene suksdorfii appears to be closely related to S. parryi but differs in its broadly winged seeds, smaller size, cespitose habit, and the prominent purple-septate hairs of the calyx, although the latter occasionally are present in S. parryi. It is very similar to, and in Idaho appears to intergrade with, another alpine species, S. sargentii, which has linear leaves and lacks the purple septa in the hairs and the broad wing on the seeds. It is similar also to S. hitchguirei; see discussion under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 20 or 26, species ca. 1500 (8 genera, 89 species in the flora).

Caryophylloideae can be characterized by the presence of sepals connate into a cup or (usually) long tube, clawed petals (often with appendages and auricles), and a lack of stipules. The largest genera in the family [Silene (incl. Lychnis), about 700 species; Dianthus, about 320 species] are in the Caryophylloideae; together with Gypsophila (about 150 species), these three genera include about three-quarters of the species found in the family. Three tribes are often differentiated on calyx venation and number of styles, with two, Caryophylleae and Sileneae, incorporating nearly all of the genera.

Caryophylloideae share the caryophyllad type of embryogeny with Alsinoideae and, as postulated by V. Bittrich (1993), the two may form a monophyletic group. Results from preliminary molecular studies by M. Nepokroeff et al. (2002) and R. D. Smissen et al. (2002) reinforce that hypothesis, but the relationships among members of the two subfamilies remain unclear.

Most of the molecular work within the subfamily has focused on Sileneae and more specifically on trying to determine whether or not Silene is monophyletic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 209. FNA vol. 5, p. 152. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
Synonyms family Caryophyllaceae subfamily Silenoideae
Name authority B. L. Robinson: Bot. Gaz. 16: 44, plate 6, figs. 9–11. (1891) Arnott: in M. Napier, Encycl. Brit. ed. 7, 5: 99. (1832)
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