Silene stellata |
Silene uralensis |
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starry campion, widow's frill |
apetalous catchfly, mountain campion, nodding campion, Spalding's campion |
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Habit | Plants perennial; taproot thick; caudex branched. | Plants perennial, cespitose or not; taproot stout. | ||||||||
Stems | several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base. |
erect, branched or simple, 5–40 cm, glabrous or pubescent. |
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Leaves | withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region; blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially. |
basal few or numerous, petiolate, blade narrowly oblanceolate, spatulate, 1–13 cm × 1–7 mm, margins ciliate, glabrous or pubescent; cauline in 1–5 pairs, sessile, much reduced distally, blade linear to lanceolate, 0.1–4 cm × 1–5 mm. |
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Inflorescences | paniculate, open, bracteate, bracteolate, branches elongate, puberulent; bracts and bracteoles linear-lanceolate, 2–15 mm. |
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Pedicels | straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous. |
slender or stout. |
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Flowers | calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm; corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent; stamens equaling petals; styles 3, longer than petals. |
erect or nodding; calyx prominently 10-veined, ovate-elliptic to broadly campanulate, inflated, 11–18 × 6–13 mm, thin and papery, veins purple or brown, lobes ovate to triangular, 2–3 mm; corolla dingy pink, purple, or red, claw shorter than to equaling calyx, limb not differentiated from claw, ovate or obovate, unlobed to 2-lobed, 1–4 mm; stamens shorter than petals; styles 5, shorter than petals. |
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Capsules | globose, opening by 3 broadly triangular teeth; carpophore 2–3 mm. |
equaling to slightly longer than calyx, opening by 10 recurved teeth; carpophore 1–2 mm. |
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Seeds | dark brown, reniform, ca. 1 mm, papillate. |
brown or sooty brown, broadly winged, round to ± angular, 1.5–2.5 mm diam. including wing. |
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2n | = (34), 48. |
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Silene stellata |
Silene uralensis |
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Phenology | Flowering summer. | |||||||||
Habitat | Rich deciduous woods, river flats, tall-grass prairies | |||||||||
Elevation | 0-1300 m (0-4300 ft) | |||||||||
Distribution |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
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AK; CO; MT; UT; WY; AB; BC; MB; NL; NT; NU; ON; QC; YT; Canada; Greenland; nw United States including Alaska; e Asia (Russian Far East, Siberia)
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Discussion | Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered. Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). Silene uralensis is a very variable species complex. Recently, A. Kurtto (2001) has resurrected S. wahlbergella as a species distinct from S. uralensis, differing in its shorter petals and larger capsule. It appears to be confined to Scandanavia and adjoining arctic Russia. North American reports of S. wahlbergella (and its synonym Lychnis apetala) are all referable to S. uralensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 207. | FNA vol. 5, p. 209. | ||||||||
Parent taxa | Caryophyllaceae > subfam. Caryophylloideae > Silene | Caryophyllaceae > subfam. Caryophylloideae > Silene | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Cucubalus stellatus, S. scabrella, S. stellata var. scabrella | Gastrolychnis uralensis | ||||||||
Name authority | (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) | (Ruprecht) Bocquet: Candollea 22: 25. (1967) | ||||||||
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