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starry campion, widow's frill

apetalous catchfly, mountain campion, nodding campion, Spalding's campion

Habit Plants perennial; taproot thick; caudex branched. Plants perennial, cespitose or not; taproot stout.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

erect, branched or simple, 5–40 cm, glabrous or pubescent.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

basal few or numerous, petiolate, blade narrowly oblanceolate, spatulate, 1–13 cm × 1–7 mm, margins ciliate, glabrous or pubescent;

cauline in 1–5 pairs, sessile, much reduced distally, blade linear to lanceolate, 0.1–4 cm × 1–5 mm.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

slender or stout.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

erect or nodding;

calyx prominently 10-veined, ovate-elliptic to broadly campanulate, inflated, 11–18 × 6–13 mm, thin and papery, veins purple or brown, lobes ovate to triangular, 2–3 mm;

corolla dingy pink, purple, or red, claw shorter than to equaling calyx, limb not differentiated from claw, ovate or obovate, unlobed to 2-lobed, 1–4 mm;

stamens shorter than petals;

styles 5, shorter than petals.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

equaling to slightly longer than calyx, opening by 10 recurved teeth;

carpophore 1–2 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

brown or sooty brown, broadly winged, round to ± angular, 1.5–2.5 mm diam. including wing.

2n

= (34), 48.

Silene stellata

Silene uralensis

Phenology Flowering summer.
Habitat Rich deciduous woods, river flats, tall-grass prairies
Elevation 0-1300 m (0-4300 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; MT; UT; WY; AB; BC; MB; NL; NT; NU; ON; QC; YT; Canada; Greenland; nw United States including Alaska; e Asia (Russian Far East, Siberia)
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Silene uralensis is a very variable species complex. Recently, A. Kurtto (2001) has resurrected S. wahlbergella as a species distinct from S. uralensis, differing in its shorter petals and larger capsule. It appears to be confined to Scandanavia and adjoining arctic Russia. North American reports of S. wahlbergella (and its synonym Lychnis apetala) are all referable to S. uralensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences usually simple, pedicels slender with single nodding (deflexed) flower (fruiting pedicels erect); seeds 1.5-2(-2.5) mm diam. (incl. broad wing).
subsp. uralensis
1. At least some of the inflorescences branched with 2 to several flowers, pedicels erect to angled but not deflexed except at tip in flower; seeds 2-2.5 mm diam
→ 2
2. Stems usually glabrous to sparsely, rarely pubescent, (15-)20-40 cm, slender; inflorescences branched with (1-)2-10 flowers; corolla only slightly exceeding calyx
subsp. ogilviensis
2. Stems densely pubescent with purple-septate hairs, 10-35 cm, stout; at least some inflorescences forked with 1-3(-4) flowers; corolla usually ca. 11/ 2 times calyx
subsp. porsildii
Source FNA vol. 5, p. 207. FNA vol. 5, p. 209.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. uralensis subsp. ogilviensis, S. uralensis subsp. porsildii, S. uralensis subsp. uralensis
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella Gastrolychnis uralensis
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) (Ruprecht) Bocquet: Candollea 22: 25. (1967)
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