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starry campion, widow's frill

creeping silene, pink campion

Habit Plants perennial; taproot thick; caudex branched. Plants perennial; taproot slender; caudex branched, woody, producing subterranean creeping stems and clumps of flowering and vegetative shoots.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

erect to straggling, branched or simple, leafy, 7–35 cm, retrorse-puberulent.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

2 per node, sessile, blade 1-veined, linear, lanceolate, or narrowly oblong, tapered to base, apex acute, puberulent on both surfaces.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

cymose, compound, ca. (2–)5–20-flowered, usually compact, bracteate;

bracts narrowly lanceolate, 2–15 mm.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

ascending, 0.1–1(–2) cm, shorter or longer than calyx, eglandular or with scattered glands, pubescence white, dense, short.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

calyx obscurely 10-veined, tubular-campanulate, 10–15 × 3–5 mm in flower, becoming clavate and 5–6 mm broad in fruit, herbaceous, villous, veins purple tinged or entirely purple, without conspicuous pale commissures, lobes ovate, 1–2.5 mm, margins broad, scarious, apex obtuse;

corolla rose pink, rarely white, clawed, claw equaling calyx, broadened distally, limb obconic, 2-lobed, 3–7 mm, appendages 0.7–1.2 mm;

stamens equaling calyx;

styles 3(–4), exserted.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

ovoid, equaling calyx, opening by 6 (or 8) spreading teeth;

carpophore 4–7 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

grayish brown, not winged, reniform, 0.8–1 mm, rugose on sides, margins shortly papillate.

2n

= (34), 48.

= 24, 48.

Silene stellata

Silene repens

Phenology Flowering summer. Flowering summer.
Habitat Rich deciduous woods, river flats, tall-grass prairies Arctic and mountain areas, dry grassy slopes, open woods, sagebrush, rocky outcrops, talus, gravel flats
Elevation 0-1300 m (0-4300 ft) 0-3200 mm
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
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[BONAP county map]
from FNA
AK; ID; MT; WA; WY; BC; NT; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Three subspecies have been recognized within Silene repens on the basis of stature and the development of purple pigment in the calyx. Northern populations in the flora area have been referred to subsp. purpurata because of the unusually heavy pigment of the calyx, while the disjunct populations in the central Rocky Mountains have been recognized as subsp. australis. However, both of these forms occur among collections from Eurasia that have been referred to subsp. repens. When material from populations of subsp. purpurata was grown farther south, it took on the appearance of subsp. australis, suggesting that the differences are under environmental influence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 207. FNA vol. 5, p. 201.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella S. purpurata, S. repens subsp. australis, S. repens var. australis, S. repens var. costata, S. repens subsp. purpurata
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) Patrin ex Persoon: Syn. Pl. 1: 500. (1805)
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