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starry campion, widow's frill

Grand Canyon campion

Habit Plants perennial; taproot thick; caudex branched. Plants perennial; taproot stout; caudex much-branched, woody.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

several, erect from decumbent base, branched, 20–40 cm, sparsely scabrid-puberulent, stipitate-glandular distally.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

2 per node;

basal shortly petiolate, blade oblanceolate, apex acute, sparsely puberulent;

cauline sessile, in 3–6 pairs, blade lanceolate to narrowly elliptic-lanceolate, 3–8 cm × 3–8 mm, apex acute, scabrid-puberulent.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

cymose, open, branches few, ascending, elongate, 5–10 cm, flowers 1–3 per branch, bracteate;

bracts lanceolate, 3–5 mm, apex acute.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

usually shorter than or to 11/2 times calyx, scabrous-puberulent, ± viscid.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

bisexual and unisexual, all flowers having bisexual flowers and staminate unisexual flowers, 6–8 mm diam.;

calyx prominently 10-veined, narrowly campanulate in flower, campanulate in fruit, 7–10(–11) × 4–6 mm, only slightly contracted around carpophore, papery and membranous, sparsely scabrid with short glandular-setose hairs, veins parallel, green, with pale commissures, margins dentate, lobes broadly ovate (as broad as long), 1–2 mm, not ciliate;

corolla cream or white, tinged purple, claw slightly exceeding calyx, narrowly triangular, auriculate, puberulent at base, limb oblong, 2-lobed, 2–4 mm, lobes entire, appendages 0.1–0.5 mm and broad, entire;

stamens equaling petals;

styles 3, equaling petals, reduced in functionally staminate flowers.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

ovoid-ellipsoid, equaling calyx, opening by 6 spreading teeth;

carpophore ca. 2 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

brown, reniform-triangular, ca. 1 mm; papillate mainly around margin.

2n

= (34), 48.

Silene stellata

Silene rectiramea

Phenology Flowering summer. Flowering April-May.
Habitat Rich deciduous woods, river flats, tall-grass prairies Pinyon pine woodland, dry slopes
Elevation 0-1300 m (0-4300 ft) 1800-2100 m (5900-6900 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
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from FNA
AZ
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Silene rectiramea is found in the Grand Canyon in Arizona. It is very similar to forms of S. verecunda and to S. thurberi. It is distinguished by its smaller flowers and fruiting calyces that are not or only slightly constricted around the carpophore and tend to be more sparsely pubescent with broader, shorter lobes. Also the appendages smaller and entire.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 207. FNA vol. 5, p. 200.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) B. L. Robinson: Bot. Gaz. 28: 134. (1899)
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