Silene stellata |
Silene parryi |
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starry campion, widow's frill |
Parry's, Parry's campion, Parry's catchfly, Parry's silene |
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Habit | Plants perennial; taproot thick; caudex branched. | Plants perennial; taproot thick; caudex branched, woody, with tufts of basal leaves. |
Stems | several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base. |
erect, simple, (10–)20–60 cm, softly puberulent, viscid-glandular distally. |
Leaves | withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region; blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially. |
mostly basal; basal petiolate, blade oblanceolate, spatulate, 3–8 cm × 2–14 mm, not fleshy, margins shortly ciliate, apex ± acute, glabrous to puberulent on both surfaces; cauline usually in 2–4 pairs, blade narrowly oblanceolate, lanceolate to linear-lanceolate, 0.2–0.8 cm × 10–80 mm, not fleshy, puberulent on both surfaces, at least distal ones glandular. |
Inflorescences | paniculate, open, bracteate, bracteolate, branches elongate, puberulent; bracts and bracteoles linear-lanceolate, 2–15 mm. |
(1–)3–7-flowered, open, bracteate; bracts linear-lanceolate, broadened at base, 2–10 mm. |
Pedicels | straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous. |
ascending, usually longer than calyx, puberulent, viscid stipitate-glandular. |
Flowers | calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm; corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent; stamens equaling petals; styles 3, longer than petals. |
calyx prominently 10-veined, campanulate, inflated, ± umbilicate, not or only slightly constricted toward base, (10–)12–16 × 7–9 mm in fruit, glandular-pubescent, strongly viscid, veins parallel, purplish, with pale commissures, not much broadened distally, commissural veins slender, forked distally and fused to those of lobes, lobes ovate to broadly triangular with lanceolate midrib, 2–3 mm, margins purple tinged, broad, membranous; corolla white, often tinged green or purple, clawed, claw equaling calyx, glabrous, broadened distally, limb deeply 2-lobed, rarely 4-lobed, 5–7 mm, lobes with 2 prominent lateral teeth, appendages 2(–4), 1.5–2 mm; stamens equaling calyx; stigmas 3(–5), exserted. |
Capsules | globose, opening by 3 broadly triangular teeth; carpophore 2–3 mm. |
included in calyx, opening by 3(–5) teeth, each tardily splitting into 2; carpophore 2–3 mm. |
Seeds | dark brown, reniform, ca. 1 mm, papillate. |
brown, not winged, broadly reniform and often flattened, 1.5–2.5 mm, rugose to shallowly tuberculate on sides, larger tubercules on margins. |
2n | = (34), 48. |
= 48, 96. |
Silene stellata |
Silene parryi |
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Phenology | Flowering summer. | Flowering summer. |
Habitat | Rich deciduous woods, river flats, tall-grass prairies | Mountains, gravelly ridges, rocky and grassy slopes, subalpine meadows, grassy openings in montane forests |
Elevation | 0-1300 m (0-4300 ft) | 1500-3000 m (4900-9800 ft) |
Distribution |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
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ID; MT; OR; WA; WY; AB; BC
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Discussion | Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered. Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Silene parryi is very similar to S. douglasii, but the latter is normally eglandular with a characteristic short, gray, retrorse pubescence. The two species may hybridize, accounting for the occurrence of populations of S. douglasii with some glandular pubescence in the inflorescence. Silene parryi is closely related also to S. scouleri, but the latter is normally readily distinguished by its pink flowers; taller stature; long, narrow, many-flowered inflorescences; and fusiform fruiting calyces that are constricted around the carpophore. However, some depauperate specimens of S. scouleri from montane habitats are difficult to place. Also, small plants of S. parryi from alpine habitats can easily be mistaken for S. grayi. The anthers of S. parryi are often smutted with Microbotryum violaceum (Persoon) G. Deml & Oberwinker [= Ustilago violacea (Persoon) Roussel], e.g., in the type collection of S. tetonensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 207. | FNA vol. 5, p. 197. |
Parent taxa | Caryophyllaceae > subfam. Caryophylloideae > Silene | Caryophyllaceae > subfam. Caryophylloideae > Silene |
Sibling taxa | ||
Synonyms | Cucubalus stellatus, S. scabrella, S. stellata var. scabrella | Lychnis parryi, S. douglasii var. macounii, S. macounii, S. scouleri var. macounii, S. tetonensis, S. tetragyna, Wahlbergella parryi |
Name authority | (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) | (S. Watson) C. L. Hitchcock & Maguire: Revis. N. Amer. Silene, 36. (1947) |
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