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starry campion, widow's frill

Drummond's campion, Drummond's catchfly, fork catchfly

Habit Plants perennial; taproot thick; caudex branched. Plants perennial; taproot stout; caudex branched, somewhat fleshy.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

erect, simple or several from base, retrorsely puberulent proximally, densely so and viscid distally, with stipitate glands.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

blade with stiff, appressed pubescence on both surfaces;

basal petiolate, blade lanceolate to elliptic or oblanceolate, (1.5–)3–10 cm × 4–12 mm (including petiole);

cauline in 2–5 pairs, blade linear to linear-lanceolate, 3–9 cm × 2–7 mm.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

1–20-flowered, bracteate, strongly viscid-glandular or less densely pubescent, longer hairs sometimes purple-septate;

bracts narrowly lanceolate, thick, 3–15 mm, herbaceous, apex acuminate.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

stiffly erect, 0.1–5 cm, varying in length within same inflorescence.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

calyx 10-veined, broadly tubular to narrowly ellipsoid, not inflated, 12–18 × 4–8 mm in fruit, 2–3 times as long as broad, membranous between veins, margins dentate with 5 triangular, 1.2–2 mm lobes erect in flower and spreading in fruit, apex acuminate, veins green;

petals off-white to dusky pink or dingy reddish purple, clawed, equaling or to 11/2 times calyx, claw broadened distally, limb not differentiated from claw, narrower than claw, 1–3 mm;

stamens included in calyx;

styles (4–)5, included in calyx.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

12–15 mm, equaling calyx (rarely to 11/2 times calyx), opening by (4–)5 spreading teeth.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

dark brown, not winged, reniform to angular, 0.7–1 mm diam., margins finely papillate;

papillae triangular, slender, longer than broad.

2n

= (34), 48.

Silene stellata

Silene drummondii

Phenology Flowering summer.
Habitat Rich deciduous woods, river flats, tall-grass prairies
Elevation 0-1300 m (0-4300 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Subspecies drummondii is characteristically a prairie taxon, while subsp. striata is associated with the Rocky Mountains. However, the two taxa frequently appear to intergrade; e.g., in the Cypress Hills of southern Alberta and Saskatchewan, and in the southern Rockies. Variety kruckebergii appears to be a luxuriant form with a more elongate capsule and calyx. Silene invisa, a Californian species, is similar to S. drummondii, some plants of which, from Nevada and Arizona, tend to be intermediate (see note under S. invisa).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals equaling calyx; fruiting calyces 12-15 × 4-6 mm, 21/ 4-3 times as long as broad; seeds ca. 0.7 mm diam.; inflorescences typically (1-)3-10(-20)-flowered
subsp. drummondii
1. Petals 11/ 1/ 2 times calyx and clearly exserted from it; fruiting calyces 13-18 × 6-8 mm, ca. 2 times as long as broad; seeds ca. 1 mm diam.; inflorescences typically 1-4(-8)-flowered
subsp. striata
Source FNA vol. 5, p. 207. FNA vol. 5, p. 183.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. drummondii subsp. drummondii, S. drummondii subsp. striata
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella Gastrolychnis drummondii, Lychnis drummondii, Melandrium drummondii, Wahlbergella drummondii
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) Hooker: Fl. Bor.-Amer. 1: 89. (1830)
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