Silene stellata
Silene dioica
starry campion, widow's frill
red campion, red catchfly, silène dioïque
several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.
ascending, decumbent at base, branched, shortly rhizomatous, to 80 cm, softly pubescent, ± glandular, at least distally, rarely subglabrous.
withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;
blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.
sessile at mid and distal stem, petiolate to spatulate proximally;
petiole equaling or longer than blade of basal leaves;
blade ovate to elliptic, 3–13 cm × 10–50 mm (not including petiole), apex acute to acuminate, sparingly pubescent, densely so on abaxial midrib.
paniculate, open, bracteate, bracteolate, branches elongate, puberulent;
bracts and bracteoles linear-lanceolate, 2–15 mm.
dichasial cymes, several- to many-flowered, open, bracteate;
bracts lanceolate, 4–20 × 2–7 mm, herbaceous, softly hairy throughout with long-septate hairs, not glandular, or with some glandular hairs.
straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.
ascending, 0.2–3 cm, usually shorter than calyx.
calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;
corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;
stamens equaling petals;
styles 3, longer than petals.
unisexual, some plants having only staminate flowers, others having only pistillate flowers, 20–25 mm diam.;
calyx 8–12-veined, campanulate, narrowly so in staminate flowers, broadly in pistillate, 10–15 × to 7 mm in flower, 11 mm broad in fruit, herbaceous, margins dentate, softly pubescent, lobes 5, erect, lanceolate, 2–3 mm;
petals bright pink, clawed, claw equaling or longer than calyx, limb spreading horizontally, broadly obovate, unlobed or 2-lobed, to 12 × 12 mm, appendages 4, ca. 1 mm;
stamens and stigmas equaling petal claw;
styles 5.
globose, opening by 3 broadly triangular teeth;
carpophore 2–3 mm.
broadly ovoid to globose, equaling and often splitting calyx, opening by 5 (splitting into 10) revolute teeth;
carpophore absent.
dark brown, reniform, ca. 1 mm, papillate.
dark brown to black, broadly reniform, plump, 1–1.6 mm, densely and evenly papillate.
= (34), 48.
= 24.
Silene stellata
Silene dioica
Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.
Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Silene dioica is closely related to S. latifolia and completely interfertile with it. The two species hybridize wherever they grow in close proximity, and the offspring (S. hampeana Meusel & K. Werner) usually have pale pink flowers. Silene dioica and S. latifolia are difficult to separate in herbarium material unless flower color has been noted. The characters that distinguish S. dioica are the usually dense, long, and soft pubescence covering at least the distal portion of the plant; the broad, almost globose, thin, and brittle capsule with revolute teeth; and the softer, thinner, usually broader leaves. Occasionally, double-flowered plants are encountered as garden escapes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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