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starry campion, widow's frill

Balkan campion, Balkan catchfly, biennial campion, silène bisannuel

Habit Plants perennial; taproot thick; caudex branched. Plants annual or biennial, glabrous and somewhat glaucous; tap-root stout.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

erect, sparingly branched below inflorescence, robust, to 65 cm.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

basal few, usually withering by time of anthesis, blade spatulate;

cauline numerous, 2 per node, blade 1-veined, ovate-lanceolate to oblanceolate, 3–7 cm × 7–30 mm, margins entire, apex acute.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

many-flowered, open, bracteate;

primary branches racemose, elongate, with sessile or shortly pedunculate cymes of 1–6 flowers per node;

bracts narrowly lanceolate, 3–12 mm, hyaline-margined, apex acute.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

ascending, ± straight, 1–2 times calyx, broadening at calyx base.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

calyx often obscurely ca. 20-veined, elliptic, abruptly contracted at base, opening constricted to 1/2 its diam., slightly inflated in flower, 7–10 × 3–4 mm, in fruit tightly enveloping capsule, ovoid, thin, enlarging to 9–13 × 5–7 mm, herbaceous, margins narrow, membranous, dentate with broadly triangular lobes to 1 mm, glabrous, veins obscure, usually purple tinged, without conspicuous, pale commissures, longitudinal, parallel, not obviously reticulate;

petals white, clawed, claw equaling calyx, limb deeply 2-fid into 2 spatulate lobes, to 5 mm, appendages ca. 0.5 mm;

stamens exserted, to 2 times length of calyx;

filaments usually dark purple;

stigmas 3, exserted, to 2 times length of calyx.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

ovoid, equaling calyx and sometimes splitting it, opening by 6 recurved, narrowly lanceolate teeth;

carpophore ca. 1 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

grayish brown, plump, broadly reniform, 0.6–1 mm, with concentric rings of papillae;

papillae slightly longer than broad.

2n

= (34), 48.

= 24.

Silene stellata

Silene csereii

Phenology Flowering summer. Flowering summer.
Habitat Rich deciduous woods, river flats, tall-grass prairies Cultivated fields, roadsides, waste land
Elevation 0-1300 m (0-4300 ft) 0-1600 m (0-5200 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NY; OH; PA; SD; VT; WA; WI; WY; AB; BC; MB; ON; QC; SK; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Often confused with Silene vulgaris, S. csereii may be readily separated by the long, racemose primary branches of its inflorescence, the elliptic calyx that is constricted at both ends, tightly enclosing the capsule and lacking obvious reticulate venation, and the purple filaments.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 207. FNA vol. 5, p. 180.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) Baumgarten: Enum. Stirp. Transsilv. 3: 345. (1816)
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