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starry campion, widow's frill

sticky catchfly, wild campion, wild pink

Habit Plants perennial; taproot thick; caudex branched. Plants perennial, cespitose; tap-root stout; caudex much-branched, woody.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

ascending, scarcely branched, 8–20(–30) cm, softly pubescent, stipitate-glandular or eglandular, rarely glabrate.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

mostly basal, petiolate, 3–12 cm (including petiole);

cauline leaves in 2–4 pairs, those of mid and distal stem sessile, shorter and narrower;

blade narrowly to broadly oblanceolate, base spatulate into winged petiole, apex acute to obtuse, glabrous, puberulent or pilose on both surfaces and frequently stipitate-glandular, at least petioles usually ciliate.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

(1–)3–15-flowered, open, bracteate;

bracts foliaceous.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

ascending or erect, 0.2–0.8(–1.5) cm, densely pubescent and frequently stipitate-glandular.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

calyx usually green, prominently 8–10-veined, in flower narrowly tubular, lobed, narrowed proximally around carpophore, 15–22 × to 5 mm, becoming broader and clavate in fruit, pilose or stipitate-glandular, veins parallel, with pale commissures, lobes round, 1–3 mm, margins usually purple tinged, broad, membranous;

petals spreading, usually bright pink, rarely white, broadly to narrowly obovate, 2 times longer than calyx, base tapered into ciliate claw equaling or slightly longer than calyx, margins entire or shallowly lobed and crenulate, auricles absent, appendages oblong, unlobed, 1.5–2 mm;

stamens equaling claw;

filaments glabrous;

styles 3(–4), ultimately slightly exceeding claw.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

ellipsoid to obovoid, 8–10 mm, equaling calyx, opening by 6 (or 8) recurved teeth;

carpophore 5–8 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

dark brown, reniform-rotund, 1.3–1.5 mm, coarsely and evenly papillate.

2n

= (34), 48.

Silene stellata

Silene caroliniana

Phenology Flowering summer.
Habitat Rich deciduous woods, river flats, tall-grass prairies
Elevation 0-1300 m (0-4300 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; DE; GA; KS; KY; MA; MD; MO; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Although the three subspecies of Silene caroliniana have overlapping ranges of distribution, subsp. caroliniana occurs predominently in the southeastern United States, subsp. pensylvanica in the northeast, and subsp. wherryi on the western side of the Appalachians. Intermediate plants are occasionally encountered. A hybrid between subsp. wherryi and S. virginica was reported by J. A. Steyermark (1963), and a hybrid swarm between subsp. pensylvanica and S. virginica by R. S. Mitchell and L. J. Uttal (1969).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyces with long, straight, nonglandular pubescence
subsp. wherryi
1. Calyces glandular-pubescent
→ 2
2. Basal leaf blades pubescent on both surfaces, typically obovate to oblanceolate, apex obtuse; petioles broadly winged
subsp. caroliniana
2. Basal leaf blades ± glabrous on both surfaces (margins and veins pubescent abaxially), narrowly oblanceolate, apex acute; petioles with very narrow wing
subsp. pensylvanica
Source FNA vol. 5, p. 207. FNA vol. 5, p. 177.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. caroliniana subsp. caroliniana, S. caroliniana subsp. pensylvanica, S. caroliniana subsp. wherryi
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) Walter: Fl. Carol., 142 [as 241]. (1788)
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