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starry campion, widow's frill

bell catchfly, campanulate campion, red mountain catchfly, slender campion

Habit Plants perennial; taproot thick; caudex branched. Plants perennial; taproot stout; caudex much-branched, woody, producing many erect-to-straggling, little-branched flowering shoots.
Stems

several, simple proximal to inflorescence, 30–80 cm, puberulent, becoming subglabrous near base.

erect, 5–40 cm, softly pubescent to scabrous, eglandular or viscid-glandular, especially distally, very rarely glabrous, with several pairs of leaves equaling or shorter than internodes.

Leaves

withering proximally, in whorls of 4, ± sessile to short-petiolate, largest in mid-stem region;

blade lanceolate to ovate-lanceolate, 3–10 cm × 4–40 mm, apex acuminate, puberulent on both surfaces, sparsely so adaxially.

sessile, or basal with short pseudopetiole;

blade linear to lanceolate or broadly ovate, base round to cuneate, apex acute to shortly acuminate, puberulent on both surfaces, sometimes glandular.

Inflorescences

paniculate, open, bracteate, bracteolate, branches elongate, puberulent;

bracts and bracteoles linear-lanceolate, 2–15 mm.

usually with single dichotomy, rarely double, open, bracteate, branches often elongate, flowers 1 per node;

bracts foliaceous.

Pedicels

straight, often with 1 or 2 pairs of bracteoles, slender, 1/2–3 times calyx, glabrous or scabrous-puberulous.

sharply reflexed at base, especially after anthesis, equaling calyx.

Flowers

calyx obscurely 10-veined, broadly campanulate, becoming obtriangular in fruit, 7–11 × 6–10 mm, herbaceous, margins dentate, very narrow, membranous, sparsely puberulent, lobes broadly triangular, 2–3 mm;

corolla white, ca. 2 times longer than calyx, limb obtriangular, narrowed into claw, divided ca. 1/2 its length into 4–12 lobes, appendages absent;

stamens equaling petals;

styles 3, longer than petals.

nodding;

calyx obscurely 10-veined, broadly campanulate, lobed, 6–8 mm, enlarging to 13–16 mm in fruit, herbaceous, usually with short, dense pubescence throughout, often glandular-viscid, veins green, rarely purplish tinged, conspicuous pale commissures absent;

lobes ovate-triangular, 1/2 to equaling tube, herbaceous;

petals creamy white, often greenish abaxially, rarely pink tinged to dusky pink (subsp. campanulata), clawed, to 2 times calyx, claw villose, limb deeply divided and fan-shaped with many narrow, linear lobes, lobes rapidly curling, margins deeply divided or erose, appendages 2–4, to 2 mm;

stamens exserted;

filaments hairy at base;

styles 3, to 2 times calyx.

Capsules

globose, opening by 3 broadly triangular teeth;

carpophore 2–3 mm.

ovoid, ca. equaling calyx and often splitting it, opening by 6 broadly triangular teeth;

carpophore 1–2.5 mm.

Seeds

dark brown, reniform, ca. 1 mm, papillate.

brown, reniform, 2–2.5 mm broad, coarsely and ± evenly papillate;

papillae ca. as long as broad.

2n

= (34), 48.

Silene stellata

Silene campanulata

Phenology Flowering summer.
Habitat Rich deciduous woods, river flats, tall-grass prairies
Elevation 0-1300 m (0-4300 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene stellata is a very distinct species with its broadly lanceolate leaves in groups of four at each node, and its brilliant white, multilobed petals. Two varieties are recognized by some workers: var. stellata, with glabrous pedicels; and var. scabrella, with scabrous pedicels. The former tends to have longer, more slender pedicels and be more common towards the northeast, whereas the latter tends to be more western. The correlation of characters and distribution is poor, however, and intermediate plants are often encountered.

Silene stellata was collected near the Grand River, Cambridge, Ontario, in 1941, but was probably introduced there and has not been seen since.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals dusky pink; leaf blades linear to lanceolate, usually less than 10 mm broad; stem pubescence scabrid, hairs 2 or 4 times as long as broad
subsp. campanulata
1. Petals creamy white; leaf blades lanceolate to ovate, usually more than 10 mm broad; stem pubescence not scabrid, hairs at least 5 times as long as broad
subsp. glandulosa
Source FNA vol. 5, p. 207. FNA vol. 5, p. 176.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. campanulata subsp. campanulata, S. campanulata subsp. glandulosa
Synonyms Cucubalus stellatus, S. scabrella, S. stellata var. scabrella
Name authority (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 3: 84. (1811) S. Watson: Proc. Amer. Acad. Arts 10: 341. (1875)
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