Silene scaposa |
Silene scouleri |
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Blue Mountain catchfly, Robinson's catchfly, stem campion |
Scouler's campion, Scouler's catchfly, Scouler's siliene, simple campion |
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Habit | Plants perennial, subscapose, cespitose; taproot stout; caudex branched, woody. | Plants perennial; taproot stout; caudex branched, woody, crowns 1–several. | ||||||||
Stems | several, erect, simple, 15–50 cm, puberulent, viscid-glandular distally. |
erect, simple proximal to inflorescence, slender or stout, 10–80 cm, puberulent. |
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Leaves | mostly basal; basal marcescent, long-petiolate, densely tufted, blade 1-veined, narrowly oblanceolate, 2–10(–20) cm × 2–12(–20) mm, not fleshy, base tapering to petiole, apex acute to obtuse, finely puberulent on both surfaces; cauline in 1–3 pairs, sessile, much reduced, blade linear-lanceolate, not fleshy. |
2 per node; basal petiolate, blade oblanceolate, 6–25 cm × 4–30 mm, retrorsely puberulent on both surfaces; cauline in 1–12 pairs, usually sessile, blade well developed, lanceolate to ovate-lanceolate, oblanceolate, or rarely linear or linear-lanceolate. |
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Inflorescences | 1–5(–7)-flowered, with terminal flower and lateral, open, pedunculate cymes often reduced to single flowers, bracteate; bracts narrowly lanceolate, 3–10(–20) mm. |
cymose, pseudo-racemose, or rarely paniculate, erect or nodding, with 1–12 flowering nodes, 2–20-flowered, open or dense, flowers paired or in many-flowered whorls, bracteate, cymes often sessile; bracts 3–60 mm. |
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Pedicels | erect, elongate, 0.5–4.5 cm, glandular-puberulent. |
becoming deflexed at base of calyx, 1/4–2 times calyx, glandular-pubescent. |
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Flowers | calyx prominently 10-veined, those to lobes lance-shaped broadened and thickened distally, commissural veins slender, not forked distally, campanulate, 10–12 × 3.5–5 mm in flower, enlarging to 15 × 10 mm in fruit, not contracted around carpophore, papery, margins dentate, glandular-pubescent, viscid, veins parallel, with pale commissures, lobes patent, ovate, 1.5–4 mm, rigid, margins broad, membranous; corolla off-white to dingy purple-red, clawed, claw exceeding calyx, ciliate proximally, broadened distally, limbs erect, 2–4-lobed, less than 1/2 length of calyx, lobes 2–5 mm, appendages 2–4, 0.5–1 mm; stamens slightly exserted; filaments lanate, expanded at base; styles 3–5, ± equaling calyx. |
shortly pedicellate or sessile; calyx prominently 10-veined, campanulate or tubular in flower, clavate, turbinate, or fusiform in fruit, constricted or not at base around carpophore in fruit, 8–20 × 3–8 mm, veins parallel, purplish or green, with pale commissures; lobes lanceolate, 2–5 mm, apex obtuse with broad membranous margin and tip; corolla white, greenish white, or pink, sometimes tinged pink or purple, clawed, claw longer than calyx, limb deeply 2–4-lobed, often with smaller lateral teeth, 2.5–8 mm, appendages 1–3 mm; stamens ± equaling corolla claw; styles 3–4, ± equaling corolla claw. |
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Capsules | slightly longer than calyx, opening by 3–5 teeth; carpophore 1.5–2.5 mm. |
ovoid to ellipsoid, equaling or slightly longer than calyx, opening by 6 or 8 teeth; carpophore 1.5–6 mm. |
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Seeds | brown, reniform, 1.2–2 mm, margins with large, inflated papillae, rugose on sides. |
brown or grayish brown, reniform, 1–1.5 mm, margins papillate, rugose on sides. |
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2n | = 48. |
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Silene scaposa |
Silene scouleri |
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Phenology | Flowering early summer. | |||||||||
Habitat | Subalpine grassy, gravelly, or rocky slopes, ponderosa pine forests, juniper scrub, sagebrush | |||||||||
Elevation | 900-3000 m (3000-9800 ft) | |||||||||
Distribution |
CO; ID; NV; OR
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AZ; CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC; Mexico
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Discussion | Silene scaposa is a very distinct species with its subscapose inflorescence, coronalike ring of short petals, and distended fruiting calyx in which the veins to the lobes are markedly broadened and lanceolate. Variation in lobing of the corolla has been the basis for recognizing two varieties: var. scaposa (var. typica C. L. Hitchcock & Maguire), which has two-lobed petals, and var. lobata, which has four-lobed petals. However, these differences appear to be of little significance. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). Silene scouleri is a very complex species that appears to be in the process of diverging into at least three different entities. Subspecies scouleri is a plant of the Pacific coast and lowlands. It has tall, stiffly erect stems, lanceolate to broadly lanceolate leaves, and a viscid inflorescence with many-flowered whorls of almost sessile flowers ranging in color from greenish white to rich pink. At the other extreme is subsp. pringlei, a plant of the mountains in Mexico extending northwards into Arizona and New Mexico. It has slender, somewhat nodding flowering stems with very narrow leaves. The flowers are usually paired at each node and secund on slender pedicels about equaling the calyx in length. The petals are off-white, sometimes tinged with dusky purple. Between the two extremes is subsp. hallii, a short, stocky plant of the Rocky Mountains and foothills with a few-flowered inflorescence. It has a larger, campanulate calyx, and some of the flowers usually become deflexed. Differentiation among these three forms is incomplete and plants indeterminate to subspecies are frequently encountered in areas away from the main distribution centers of the three subspecies. In northern Oregon and Idaho there appear to be populations connecting S. scouleri with S. oregana. They have some of the characteristics of S. oregana but not its laciniate petals. They may represent a more luxuriant form growing in taller vegetation, but their status needs further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 202. | FNA vol. 5, p. 204. | ||||||||
Parent taxa | Caryophyllaceae > subfam. Caryophylloideae > Silene | Caryophyllaceae > subfam. Caryophylloideae > Silene | ||||||||
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Synonyms | S. scaposa var. lobata | |||||||||
Name authority | B. L. Robinson: Proc. Amer. Acad. Arts 28: 145. (1893) | Hooker: Fl. Bor.-Amer. 1: 88. (1830) | ||||||||
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