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Blue Mountain catchfly, Robinson's catchfly, stem campion

Scouler's campion, Scouler's catchfly, Scouler's siliene, simple campion

Habit Plants perennial, subscapose, cespitose; taproot stout; caudex branched, woody. Plants perennial; taproot stout; caudex branched, woody, crowns 1–several.
Stems

several, erect, simple, 15–50 cm, puberulent, viscid-glandular distally.

erect, simple proximal to inflorescence, slender or stout, 10–80 cm, puberulent.

Leaves

mostly basal;

basal marcescent, long-petiolate, densely tufted, blade 1-veined, narrowly oblanceolate, 2–10(–20) cm × 2–12(–20) mm, not fleshy, base tapering to petiole, apex acute to obtuse, finely puberulent on both surfaces;

cauline in 1–3 pairs, sessile, much reduced, blade linear-lanceolate, not fleshy.

2 per node;

basal petiolate, blade oblanceolate, 6–25 cm × 4–30 mm, retrorsely puberulent on both surfaces;

cauline in 1–12 pairs, usually sessile, blade well developed, lanceolate to ovate-lanceolate, oblanceolate, or rarely linear or linear-lanceolate.

Inflorescences

1–5(–7)-flowered, with terminal flower and lateral, open, pedunculate cymes often reduced to single flowers, bracteate;

bracts narrowly lanceolate, 3–10(–20) mm.

cymose, pseudo-racemose, or rarely paniculate, erect or nodding, with 1–12 flowering nodes, 2–20-flowered, open or dense, flowers paired or in many-flowered whorls, bracteate, cymes often sessile;

bracts 3–60 mm.

Pedicels

erect, elongate, 0.5–4.5 cm, glandular-puberulent.

becoming deflexed at base of calyx, 1/4–2 times calyx, glandular-pubescent.

Flowers

calyx prominently 10-veined, those to lobes lance-shaped broadened and thickened distally, commissural veins slender, not forked distally, campanulate, 10–12 × 3.5–5 mm in flower, enlarging to 15 × 10 mm in fruit, not contracted around carpophore, papery, margins dentate, glandular-pubescent, viscid, veins parallel, with pale commissures, lobes patent, ovate, 1.5–4 mm, rigid, margins broad, membranous;

corolla off-white to dingy purple-red, clawed, claw exceeding calyx, ciliate proximally, broadened distally, limbs erect, 2–4-lobed, less than 1/2 length of calyx, lobes 2–5 mm, appendages 2–4, 0.5–1 mm;

stamens slightly exserted;

filaments lanate, expanded at base;

styles 3–5, ± equaling calyx.

shortly pedicellate or sessile;

calyx prominently 10-veined, campanulate or tubular in flower, clavate, turbinate, or fusiform in fruit, constricted or not at base around carpophore in fruit, 8–20 × 3–8 mm, veins parallel, purplish or green, with pale commissures;

lobes lanceolate, 2–5 mm, apex obtuse with broad membranous margin and tip;

corolla white, greenish white, or pink, sometimes tinged pink or purple, clawed, claw longer than calyx, limb deeply 2–4-lobed, often with smaller lateral teeth, 2.5–8 mm, appendages 1–3 mm;

stamens ± equaling corolla claw;

styles 3–4, ± equaling corolla claw.

Capsules

slightly longer than calyx, opening by 3–5 teeth;

carpophore 1.5–2.5 mm.

ovoid to ellipsoid, equaling or slightly longer than calyx, opening by 6 or 8 teeth;

carpophore 1.5–6 mm.

Seeds

brown, reniform, 1.2–2 mm, margins with large, inflated papillae, rugose on sides.

brown or grayish brown, reniform, 1–1.5 mm, margins papillate, rugose on sides.

2n

= 48.

Silene scaposa

Silene scouleri

Phenology Flowering early summer.
Habitat Subalpine grassy, gravelly, or rocky slopes, ponderosa pine forests, juniper scrub, sagebrush
Elevation 900-3000 m (3000-9800 ft)
Distribution
from FNA
CO; ID; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Silene scaposa is a very distinct species with its subscapose inflorescence, coronalike ring of short petals, and distended fruiting calyx in which the veins to the lobes are markedly broadened and lanceolate. Variation in lobing of the corolla has been the basis for recognizing two varieties: var. scaposa (var. typica C. L. Hitchcock & Maguire), which has two-lobed petals, and var. lobata, which has four-lobed petals. However, these differences appear to be of little significance.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Silene scouleri is a very complex species that appears to be in the process of diverging into at least three different entities. Subspecies scouleri is a plant of the Pacific coast and lowlands. It has tall, stiffly erect stems, lanceolate to broadly lanceolate leaves, and a viscid inflorescence with many-flowered whorls of almost sessile flowers ranging in color from greenish white to rich pink. At the other extreme is subsp. pringlei, a plant of the mountains in Mexico extending northwards into Arizona and New Mexico. It has slender, somewhat nodding flowering stems with very narrow leaves. The flowers are usually paired at each node and secund on slender pedicels about equaling the calyx in length. The petals are off-white, sometimes tinged with dusky purple. Between the two extremes is subsp. hallii, a short, stocky plant of the Rocky Mountains and foothills with a few-flowered inflorescence. It has a larger, campanulate calyx, and some of the flowers usually become deflexed. Differentiation among these three forms is incomplete and plants indeterminate to subspecies are frequently encountered in areas away from the main distribution centers of the three subspecies. In northern Oregon and Idaho there appear to be populations connecting S. scouleri with S. oregana. They have some of the characteristics of S. oregana but not its laciniate petals. They may represent a more luxuriant form growing in taller vegetation, but their status needs further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyces campanulate, not or only slightly clavate in fruit, 13-18(-20) × (5-)6-8 mm; inflorescences with (1-)3-6(-8) flowering nodes; plants 10-40cm; pedicels stout
subsp. hallii
1. Calyces tubular to narrowly clavate in flower, clavate, turbinate, or fusiform in fruit, (8-)10-16 × 3.5-7 mm; inflorescences with 3-12 flowering nodes; plants 20-80 cm; pedicels slender
→ 2
2. Inflorescences erect, flowers in dense pseudowhorls of usually sessile, (2-)5-20-flowered cymes, both sessile and pedicellate flowers in each cyme; pedicels erect
subsp. scouleri
2. Inflorescences nodding, flowers usually paired at each node, all pedicellate; pedicels ± equaling calyx, often sharply deflexed at baseof calyx
subsp. pringlei
Source FNA vol. 5, p. 202. FNA vol. 5, p. 204.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. scouleri subsp. hallii, S. scouleri subsp. pringlei, S. scouleri subsp. scouleri
Synonyms S. scaposa var. lobata
Name authority B. L. Robinson: Proc. Amer. Acad. Arts 28: 145. (1893) Hooker: Fl. Bor.-Amer. 1: 88. (1830)
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