The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Blue Mountain catchfly, Robinson's catchfly, stem campion

Habit Plants perennial, subscapose, cespitose; taproot stout; caudex branched, woody. Herbs, annual, biennial, or perennial; taprooted or rhizomatous, sometimes stoloniferous.
Stems

several, erect, simple, 15–50 cm, puberulent, viscid-glandular distally.

erect or ascending, seldom sprawling, decumbent, or prostrate, simple or branched.

Leaves

mostly basal;

basal marcescent, long-petiolate, densely tufted, blade 1-veined, narrowly oblanceolate, 2–10(–20) cm × 2–12(–20) mm, not fleshy, base tapering to petiole, apex acute to obtuse, finely puberulent on both surfaces;

cauline in 1–3 pairs, sessile, much reduced, blade linear-lanceolate, not fleshy.

opposite, rarely whorled, connate proximally, petiolate (basal leaves) or often sessile, not stipulate;

blade linear or subulate to ovate, not succulent or rarely so (Silene).

Inflorescences

1–5(–7)-flowered, with terminal flower and lateral, open, pedunculate cymes often reduced to single flowers, bracteate;

bracts narrowly lanceolate, 3–10(–20) mm.

terminal cymes, thyrses, fascicles, or capitula, or flowers solitary, axillary;

bracts foliaceous, scarious, or absent;

involucel bracteoles present or often absent.

Pedicels

erect, elongate, 0.5–4.5 cm, glandular-puberulent.

present or rarely flowers sessile or subsessile.

Flowers

calyx prominently 10-veined, those to lobes lance-shaped broadened and thickened distally, commissural veins slender, not forked distally, campanulate, 10–12 × 3.5–5 mm in flower, enlarging to 15 × 10 mm in fruit, not contracted around carpophore, papery, margins dentate, glandular-pubescent, viscid, veins parallel, with pale commissures, lobes patent, ovate, 1.5–4 mm, rigid, margins broad, membranous;

corolla off-white to dingy purple-red, clawed, claw exceeding calyx, ciliate proximally, broadened distally, limbs erect, 2–4-lobed, less than 1/2 length of calyx, lobes 2–5 mm, appendages 2–4, 0.5–1 mm;

stamens slightly exserted;

filaments lanate, expanded at base;

styles 3–5, ± equaling calyx.

bisexual or seldom unisexual (the species then often dioecious), often conspicuous;

perianth and androecium hypogynous;

sepals 5, connate (1/4–)1/2+ their lengths into cup or tube, (1–)5–40(–62) mm, apex not hooded or awned;

petals absent or 5, often showy, white to pink or red, usually clawed, auricles absent or sometimes present, coronal appendages sometimes present, blade apex entire or emarginate to 2-fid, sometimes dentate to lacinate;

stamens (5 or) 10 (absent in pistillate flowers), in 1 or 2 whorls, arising from base of ovary;

staminodes absent or rarely 1–10;

ovary 1-locular, sometimes 2-locular proximally (Vaccaria), or 3–5-locular (some Silene);

styles 2–3(–5) (absent in staminate flowers), distinct;

stigmas 2–3(–5) (absent in staminate flowers).

Fruits

capsules, opening by 4–6(–10) valves or teeth;

carpophore usually present.

Capsules

slightly longer than calyx, opening by 3–5 teeth;

carpophore 1.5–2.5 mm.

Seeds

brown, reniform, 1.2–2 mm, margins with large, inflated papillae, rugose on sides.

4–150(–500+), reddish to gray or often brown or black, usually reniform and laterally compressed to globose, sometimes oblong or shield-shaped and dorsiventrally compressed;

embryo peripheral and curved, or central and straight.

x

= 7, 10, 12, [13?,] 14, 15, 17, [18].

2n

= 48.

Silene scaposa

Caryophyllaceae subfam. caryophylloideae

Phenology Flowering early summer.
Habitat Subalpine grassy, gravelly, or rocky slopes, ponderosa pine forests, juniper scrub, sagebrush
Elevation 900-3000 m (3000-9800 ft)
Distribution
from FNA
CO; ID; NV; OR
[WildflowerSearch map]
[BONAP county map]
North-temperate regions; Europe (esp Mediterranean region); Asia (esp Mediterranean region e to c Asia); Africa (Mediterranean region, Republic of South Africa)
Discussion

Silene scaposa is a very distinct species with its subscapose inflorescence, coronalike ring of short petals, and distended fruiting calyx in which the veins to the lobes are markedly broadened and lanceolate. Variation in lobing of the corolla has been the basis for recognizing two varieties: var. scaposa (var. typica C. L. Hitchcock & Maguire), which has two-lobed petals, and var. lobata, which has four-lobed petals. However, these differences appear to be of little significance.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 20 or 26, species ca. 1500 (8 genera, 89 species in the flora).

Caryophylloideae can be characterized by the presence of sepals connate into a cup or (usually) long tube, clawed petals (often with appendages and auricles), and a lack of stipules. The largest genera in the family [Silene (incl. Lychnis), about 700 species; Dianthus, about 320 species] are in the Caryophylloideae; together with Gypsophila (about 150 species), these three genera include about three-quarters of the species found in the family. Three tribes are often differentiated on calyx venation and number of styles, with two, Caryophylleae and Sileneae, incorporating nearly all of the genera.

Caryophylloideae share the caryophyllad type of embryogeny with Alsinoideae and, as postulated by V. Bittrich (1993), the two may form a monophyletic group. Results from preliminary molecular studies by M. Nepokroeff et al. (2002) and R. D. Smissen et al. (2002) reinforce that hypothesis, but the relationships among members of the two subfamilies remain unclear.

Most of the molecular work within the subfamily has focused on Sileneae and more specifically on trying to determine whether or not Silene is monophyletic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 202. FNA vol. 5, p. 152. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
Synonyms S. scaposa var. lobata family Caryophyllaceae subfamily Silenoideae
Name authority B. L. Robinson: Proc. Amer. Acad. Arts 28: 145. (1893) Arnott: in M. Napier, Encycl. Brit. ed. 7, 5: 99. (1832)
Web links