Silene gallica |
Silene scouleri |
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common catchfly, silène de france, small-flower catchfly, windmill campion, windmill pink |
Scouler's campion, Scouler's catchfly, Scouler's siliene, simple campion |
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Habit | Plants annual; taproot slender. | Plants perennial; taproot stout; caudex branched, woody, crowns 1–several. | ||||||||
Stems | erect, branched, rarely simple, 15–45 cm, with long, often crinkled hairs mixed with short pubescence, viscid-glandular distally. |
erect, simple proximal to inflorescence, slender or stout, 10–80 cm, puberulent. |
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Leaves | 2 per node, blade with coarse, ascending, scabrous pubescence on both surfaces; basal few, withering, blade oblanceolate to spatulate-petiolate, 0.5–5 cm × 3–15 mm; cauline blades oblanceolate to lanceolate, 1–7 cm × 1–15 mm, apex obtuse or shortly acuminate to acute. |
2 per node; basal petiolate, blade oblanceolate, 6–25 cm × 4–30 mm, retrorsely puberulent on both surfaces; cauline in 1–12 pairs, usually sessile, blade well developed, lanceolate to ovate-lanceolate, oblanceolate, or rarely linear or linear-lanceolate. |
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Inflorescences | open, with racemose branches, internodes and bracts usually ca. equaling fruiting calyx, 1–5 mm, longer proximally. |
cymose, pseudo-racemose, or rarely paniculate, erect or nodding, with 1–12 flowering nodes, 2–20-flowered, open or dense, flowers paired or in many-flowered whorls, bracteate, cymes often sessile; bracts 3–60 mm. |
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Pedicels | becoming deflexed at base of calyx, 1/4–2 times calyx, glandular-pubescent. |
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Flowers | 5–8 mm diam.; calyx prominently 10-veined, narrowly tubular-ovoid in flower, ovoid in fruit, constricted at mouth, 7–10 × 3–5 mm, membranous between veins, margins dentate, hispid, hairs ca. 2 mm, veins parallel, lobes lanceolate, 2–2.5 mm, apex greenish purple, acute; petals white or pink, often with dark spot or dark pink throughout, clawed, claw equaling calyx, limb elliptic to obovate, lobed or unlobed, to 6 mm, appendages 2, oblong to narrowly lanceolate, 1–1.5 mm; stamens equaling or shorter than calyx; stigmas 3, included in calyx. |
shortly pedicellate or sessile; calyx prominently 10-veined, campanulate or tubular in flower, clavate, turbinate, or fusiform in fruit, constricted or not at base around carpophore in fruit, 8–20 × 3–8 mm, veins parallel, purplish or green, with pale commissures; lobes lanceolate, 2–5 mm, apex obtuse with broad membranous margin and tip; corolla white, greenish white, or pink, sometimes tinged pink or purple, clawed, claw longer than calyx, limb deeply 2–4-lobed, often with smaller lateral teeth, 2.5–8 mm, appendages 1–3 mm; stamens ± equaling corolla claw; styles 3–4, ± equaling corolla claw. |
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Capsules | equaling calyx, opening with 6 recurved, narrowly triangular teeth; carpophore shorter than 1 mm, pubescent. |
ovoid to ellipsoid, equaling or slightly longer than calyx, opening by 6 or 8 teeth; carpophore 1.5–6 mm. |
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Seeds | dark reddish brown, reniform, angular with concave, radially ridged faces, broad outer edge transversely ridged and verrucose, ca. 0.5 mm broad. |
brown or grayish brown, reniform, 1–1.5 mm, margins papillate, rugose on sides. |
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2n | = 24. |
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Silene gallica |
Silene scouleri |
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Phenology | Flowering spring–early summer. | |||||||||
Habitat | Dry, open places, sandy and gravelly ground, roadsides, waste land | |||||||||
Elevation | 0-2000 m (0-6600 ft) | |||||||||
Distribution |
AK; AL; AZ; CA; FL; ID; LA; MA; ME; MO; MS; NC; NH; NY; OR; PA; RI; SC; TX; WA; BC; NB; NS; ON; PE; Europe [Introduced in North America; introduced worldwide]
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AZ; CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC; Mexico
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Discussion | Subspecies 3 (3 in the flora). Silene scouleri is a very complex species that appears to be in the process of diverging into at least three different entities. Subspecies scouleri is a plant of the Pacific coast and lowlands. It has tall, stiffly erect stems, lanceolate to broadly lanceolate leaves, and a viscid inflorescence with many-flowered whorls of almost sessile flowers ranging in color from greenish white to rich pink. At the other extreme is subsp. pringlei, a plant of the mountains in Mexico extending northwards into Arizona and New Mexico. It has slender, somewhat nodding flowering stems with very narrow leaves. The flowers are usually paired at each node and secund on slender pedicels about equaling the calyx in length. The petals are off-white, sometimes tinged with dusky purple. Between the two extremes is subsp. hallii, a short, stocky plant of the Rocky Mountains and foothills with a few-flowered inflorescence. It has a larger, campanulate calyx, and some of the flowers usually become deflexed. Differentiation among these three forms is incomplete and plants indeterminate to subspecies are frequently encountered in areas away from the main distribution centers of the three subspecies. In northern Oregon and Idaho there appear to be populations connecting S. scouleri with S. oregana. They have some of the characteristics of S. oregana but not its laciniate petals. They may represent a more luxuriant form growing in taller vegetation, but their status needs further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 185. | FNA vol. 5, p. 204. | ||||||||
Parent taxa | ||||||||||
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Synonyms | S. anglica, S. quinquevulnera | |||||||||
Name authority | Linnaeus: Sp. Pl. 1: 417. (1753) | Hooker: Fl. Bor.-Amer. 1: 88. (1830) | ||||||||
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