Silene dioica
Silene occidentalis
red campion, red catchfly, silène dioïque
western campion, western catchfly
ascending, decumbent at base, branched, shortly rhizomatous, to 80 cm, softly pubescent, ± glandular, at least distally, rarely subglabrous.
erect, simple proximal to inflorescence, 30–60 cm, softly pubescent and stipitate-glandular.
sessile at mid and distal stem, petiolate to spatulate proximally;
petiole equaling or longer than blade of basal leaves;
blade ovate to elliptic, 3–13 cm × 10–50 mm (not including petiole), apex acute to acuminate, sparingly pubescent, densely so on abaxial midrib.
2 per node;
basal ± petiolate, petiole ciliate, blade oblanceolate, spatulate, 5–12 cm × 7–20 mm, apex acute, short-pubescent on both surfaces;
cauline in 3(–4) pairs, reduced distally, blade oblanceolate to lanceolate.
dichasial cymes, several- to many-flowered, open, bracteate;
bracts lanceolate, 4–20 × 2–7 mm, herbaceous, softly hairy throughout with long-septate hairs, not glandular, or with some glandular hairs.
open, narrow, with ascending branches, 9–25-flowered, bracteate, pubescent and stipitate-glandular;
bracts narrowly lanceolate, ciliate.
ascending, 0.2–3 cm, usually shorter than calyx.
1/2–3 times longer than calyx.
unisexual, some plants having only staminate flowers, others having only pistillate flowers, 20–25 mm diam.;
calyx 8–12-veined, campanulate, narrowly so in staminate flowers, broadly in pistillate, 10–15 × to 7 mm in flower, 11 mm broad in fruit, herbaceous, margins dentate, softly pubescent, lobes 5, erect, lanceolate, 2–3 mm;
petals bright pink, clawed, claw equaling or longer than calyx, limb spreading horizontally, broadly obovate, unlobed or 2-lobed, to 12 × 12 mm, appendages 4, ca. 1 mm;
stamens and stigmas equaling petal claw;
styles 5.
calyx prominently 10-veined, tubular in flower and fruit, umbilicate, somewhat constricted around carpophore, 15–38 × 3–6 mm, papery, sparsely pubescent and stipitate-glandular, veins parallel, green, with pale commissures, lobes 5, broadly ovate, 2–4 mm, scarious around green midrib;
corolla pink or rose red (rarely white), clawed, claw equaling calyx, limb oblong, fanlike, deeply 4-lobed, lobes divergent, lanceolate (rarely with only 2 lobes, each with small lateral tooth), (7–)10–20 mm, appendages linear, 2–4 mm;
stamens exserted, shorter than petals;
stigmas 3, shorter than petals.
broadly ovoid to globose, equaling and often splitting calyx, opening by 5 (splitting into 10) revolute teeth;
carpophore absent.
narrowly ovate-elliptic, longer than calyx, opening by 6 recurved teeth;
carpophore 4–18 mm.
dark brown to black, broadly reniform, plump, 1–1.6 mm, densely and evenly papillate.
grayish brown, reniform, 1–1.5 mm, verrucate.
= 24.
= 48.
Silene dioica
Silene occidentalis
Silene dioica is closely related to S. latifolia and completely interfertile with it. The two species hybridize wherever they grow in close proximity, and the offspring (S. hampeana Meusel & K. Werner) usually have pale pink flowers. Silene dioica and S. latifolia are difficult to separate in herbarium material unless flower color has been noted. The characters that distinguish S. dioica are the usually dense, long, and soft pubescence covering at least the distal portion of the plant; the broad, almost globose, thin, and brittle capsule with revolute teeth; and the softer, thinner, usually broader leaves. Occasionally, double-flowered plants are encountered as garden escapes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The long, tubular calyx and the proportionally long carpophore of Silene occidentalis are remarkable. Plants with the longest calyx tubes (more than 30 mm) have been referred to subsp. longistipitata and appear to be confined to Butte County. Calyx length varies greatly, however, and it is doubtful whether it is a justifiable basis for taxonomic recognition. The deeply lobed pink petals, together with its habit, give S. occidentalis a superficial resemblance to the European S. flos-cuculi, which occurs as an introduction on both sides of the North American continent.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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