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Balkan campion, Balkan catchfly, biennial campion, silène bisannuel

Thurber's catchfly, woolly catchfly

Habit Plants annual or biennial, glabrous and somewhat glaucous; tap-root stout. Plants perennial; taproot stout; caudex branched, woody.
Stems

erect, sparingly branched below inflorescence, robust, to 65 cm.

several, erect, freely branched, leafy, elongate, 30–80 cm, scabrid-puberulent and glandular-viscid.

Leaves

basal few, usually withering by time of anthesis, blade spatulate;

cauline numerous, 2 per node, blade 1-veined, ovate-lanceolate to oblanceolate, 3–7 cm × 7–30 mm, margins entire, apex acute.

2 per node;

basal long-petiolate, blade oblanceolate, (2–)5–18 cm × 5–30 mm, base narrowed into petiole, apex ± acute, glandular-puberulent on both surfaces;

cauline not greatly reduced in distal stem, blade 5–15 cm × 10–20 mm, viscid glandular-pubescent, sparsely so in shade forms, proximal petiolate and blade oblanceolate, distal sessile and blade lanceolate, apex acute.

Inflorescences

many-flowered, open, bracteate;

primary branches racemose, elongate, with sessile or shortly pedunculate cymes of 1–6 flowers per node;

bracts narrowly lanceolate, 3–12 mm, hyaline-margined, apex acute.

open, elongate, dichotomously branched, branches ascending, bracteate;

bracts leaflike, reduced distally.

Pedicels

ascending, ± straight, 1–2 times calyx, broadening at calyx base.

ascending, may be partially deflexed, rather slender, longer than calyx, viscid glandular-pubescent with septate hairs, septa colorless.

Flowers

calyx often obscurely ca. 20-veined, elliptic, abruptly contracted at base, opening constricted to 1/2 its diam., slightly inflated in flower, 7–10 × 3–4 mm, in fruit tightly enveloping capsule, ovoid, thin, enlarging to 9–13 × 5–7 mm, herbaceous, margins narrow, membranous, dentate with broadly triangular lobes to 1 mm, glabrous, veins obscure, usually purple tinged, without conspicuous, pale commissures, longitudinal, parallel, not obviously reticulate;

petals white, clawed, claw equaling calyx, limb deeply 2-fid into 2 spatulate lobes, to 5 mm, appendages ca. 0.5 mm;

stamens exserted, to 2 times length of calyx;

filaments usually dark purple;

stigmas 3, exserted, to 2 times length of calyx.

calyx prominently 10-veined, tubular in flower, 8–12 × 2–4 mm, swelling to campanulate and 5–7 mm broad in fruit, not contracted proximally around carpophore, viscid glandular-pubescent, especially on veins, veins parallel, green, with pale commissures, those to lobes broadened distally, lobes erect, narrowly lanceolate, 3–4 mm, rigid, setose-scabrous, glandular-viscid, apex recurved;

corolla greenish white, claw equaling calyx, limb 2-lobed, ca. 3 mm, appendages ca. 0.5 mm, margins erose;

stamens equaling petals;

styles 3, equaling petals.

Capsules

ovoid, equaling calyx and sometimes splitting it, opening by 6 recurved, narrowly lanceolate teeth;

carpophore ca. 1 mm.

slightly exserted from calyx, narrowly ovoid, opening by 6 teeth;

carpophore 1–2 mm.

Seeds

grayish brown, plump, broadly reniform, 0.6–1 mm, with concentric rings of papillae;

papillae slightly longer than broad.

almost black, ellipsoid-reniform, ca. 1 mm, coarsely papillate;

papillae inflated.

2n

= 24.

= 48.

Silene csereii

Silene thurberi

Phenology Flowering summer. Flowering late summer–fall.
Habitat Cultivated fields, roadsides, waste land Open rocky places and canyons
Elevation 0-1600 m (0-5200 ft) 1500-2000 m (4900-6600 ft)
Distribution
from FNA
CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NY; OH; PA; SD; VT; WA; WI; WY; AB; BC; MB; ON; QC; SK; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; Mexico
[BONAP county map]
Discussion

Often confused with Silene vulgaris, S. csereii may be readily separated by the long, racemose primary branches of its inflorescence, the elliptic calyx that is constricted at both ends, tightly enclosing the capsule and lacking obvious reticulate venation, and the purple filaments.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene thurberi is a rare, coarse, scabrous, and viscid herb with small, inconspicuous flowers on elongate pseudoracemes or open dichasia. The veins on the small calyces are usually green and conspicuously broaden into the recurved teeth. It is more common in Mexico.

Silene rectiramea is very similar but differs in its somewhat smaller calyx; short, ovate calyx lobes; and entire corolla appendages.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 180. FNA vol. 5, p. 209.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Synonyms S. plicata
Name authority Baumgarten: Enum. Stirp. Transsilv. 3: 345. (1816) S. Watson: Proc. Amer. Acad. Arts 10: 343. (1875)
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