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Balkan campion, Balkan catchfly, biennial campion, silène bisannuel

Sargent's campion, Sargent's catchfly

Habit Plants annual or biennial, glabrous and somewhat glaucous; tap-root stout. Plants perennial; taproot stout; caudex much-branched, woody, producing many decumbent, leafy, short shoots and erect flowering shoots, often densely cespitose.
Stems

erect, sparingly branched below inflorescence, robust, to 65 cm.

erect, decumbent at base, simple, branched in flowering region, slender, 10–20(–25) cm, sparsely pubescent and glandular.

Leaves

basal few, usually withering by time of anthesis, blade spatulate;

cauline numerous, 2 per node, blade 1-veined, ovate-lanceolate to oblanceolate, 3–7 cm × 7–30 mm, margins entire, apex acute.

mostly basal;

basal marcescent, densely tufted, long-petiolate, blade linear-oblanceolate, 1–3 cm × 0.5–3 mm, somewhat fleshy, setose-puberulent on both surfaces;

cauline in 1–3 pairs proximal to inflorescence, reduced distally, blade linear, 1–4 cm × 0.5–2 mm.

Inflorescences

many-flowered, open, bracteate;

primary branches racemose, elongate, with sessile or shortly pedunculate cymes of 1–6 flowers per node;

bracts narrowly lanceolate, 3–12 mm, hyaline-margined, apex acute.

1–3-flowered, with terminal flower, usually with 1 or 2 flowers at proximal nodes.

Pedicels

ascending, ± straight, 1–2 times calyx, broadening at calyx base.

erect, sometimes slightly bent at apex, 1/4–2 times calyx, viscid glandular-puberulent, septa of hairs colorless.

Flowers

calyx often obscurely ca. 20-veined, elliptic, abruptly contracted at base, opening constricted to 1/2 its diam., slightly inflated in flower, 7–10 × 3–4 mm, in fruit tightly enveloping capsule, ovoid, thin, enlarging to 9–13 × 5–7 mm, herbaceous, margins narrow, membranous, dentate with broadly triangular lobes to 1 mm, glabrous, veins obscure, usually purple tinged, without conspicuous, pale commissures, longitudinal, parallel, not obviously reticulate;

petals white, clawed, claw equaling calyx, limb deeply 2-fid into 2 spatulate lobes, to 5 mm, appendages ca. 0.5 mm;

stamens exserted, to 2 times length of calyx;

filaments usually dark purple;

stigmas 3, exserted, to 2 times length of calyx.

calyx prominently 10-veined, narrowly campanulate, ± umbilicate, not contracted proximally around carpophore, 9–18(–22) × 3–6 mm, papery, glandular-puberulent, viscid, hairs with colorless septa, veins parallel, purple, with pale commissures, lobes with midrib present, triangular, 2–3 mm, margins purple tinged, membranous, often broadened distally into round, crenulate lobe;

corolla off-white, usually pink or purple tinged, to 11/2 times longer than calyx, limb 2-lobed, 2–3 mm, sometimes with 2 small lateral teeth, appendages 2, ovate, 1–1.5 mm;

stamens slightly longer than calyx;

styles 3(–5), equaling petals.

Capsules

ovoid, equaling calyx and sometimes splitting it, opening by 6 recurved, narrowly lanceolate teeth;

carpophore ca. 1 mm.

included in calyx, opening by 6 (or 8 or 10) ascending teeth;

carpophore 1.5–3 mm, woolly.

Seeds

grayish brown, plump, broadly reniform, 0.6–1 mm, with concentric rings of papillae;

papillae slightly longer than broad.

brown, reniform, ca. 1.5 mm, rugose on both surfaces, margins with large, inflated papillae.

2n

= 24.

= 48.

Silene csereii

Silene sargentii

Phenology Flowering summer. Flowering summer.
Habitat Cultivated fields, roadsides, waste land Alpine grassy, gravelly, or rocky slopes and ridges, openings in subalpine forests, sagebrush, and on juniper slopes
Elevation 0-1600 m (0-5200 ft) 2400-3800 m (7900-12500 ft)
Distribution
from FNA
CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NY; OH; PA; SD; VT; WA; WI; WY; AB; BC; MB; ON; QC; SK; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Often confused with Silene vulgaris, S. csereii may be readily separated by the long, racemose primary branches of its inflorescence, the elliptic calyx that is constricted at both ends, tightly enclosing the capsule and lacking obvious reticulate venation, and the purple filaments.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene sargentii is very similar to S. suksdorfii, but that species has purple-septate hairs on the calyx and pedicels, whereas those on S. sargentii are colorless (see M. A. T. Showers 1987). Also, the cauline leaves of S. suksdorfii are narrowly oblanceolate rather than linear, and the basal leaves are marcescent. Silene sargentii may be confused with S. bernardina and can intergrade with it in Nevada. However, that species is usually larger with a longer, tubular calyx and petals that are deeply divided into 4–6 narrow lobes, unlike S. sargentii, which has 2-lobed petals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 180. FNA vol. 5, p. 202.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Synonyms Lychnis californica, S. lacustris, S. watsonii
Name authority Baumgarten: Enum. Stirp. Transsilv. 3: 345. (1816) S. Watson: Proc. Amer. Acad. Arts 14: 290. (1879)
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