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Balkan campion, Balkan catchfly, biennial campion, silène bisannuel

Grand Canyon campion

Habit Plants annual or biennial, glabrous and somewhat glaucous; tap-root stout. Plants perennial; taproot stout; caudex much-branched, woody.
Stems

erect, sparingly branched below inflorescence, robust, to 65 cm.

several, erect from decumbent base, branched, 20–40 cm, sparsely scabrid-puberulent, stipitate-glandular distally.

Leaves

basal few, usually withering by time of anthesis, blade spatulate;

cauline numerous, 2 per node, blade 1-veined, ovate-lanceolate to oblanceolate, 3–7 cm × 7–30 mm, margins entire, apex acute.

2 per node;

basal shortly petiolate, blade oblanceolate, apex acute, sparsely puberulent;

cauline sessile, in 3–6 pairs, blade lanceolate to narrowly elliptic-lanceolate, 3–8 cm × 3–8 mm, apex acute, scabrid-puberulent.

Inflorescences

many-flowered, open, bracteate;

primary branches racemose, elongate, with sessile or shortly pedunculate cymes of 1–6 flowers per node;

bracts narrowly lanceolate, 3–12 mm, hyaline-margined, apex acute.

cymose, open, branches few, ascending, elongate, 5–10 cm, flowers 1–3 per branch, bracteate;

bracts lanceolate, 3–5 mm, apex acute.

Pedicels

ascending, ± straight, 1–2 times calyx, broadening at calyx base.

usually shorter than or to 11/2 times calyx, scabrous-puberulent, ± viscid.

Flowers

calyx often obscurely ca. 20-veined, elliptic, abruptly contracted at base, opening constricted to 1/2 its diam., slightly inflated in flower, 7–10 × 3–4 mm, in fruit tightly enveloping capsule, ovoid, thin, enlarging to 9–13 × 5–7 mm, herbaceous, margins narrow, membranous, dentate with broadly triangular lobes to 1 mm, glabrous, veins obscure, usually purple tinged, without conspicuous, pale commissures, longitudinal, parallel, not obviously reticulate;

petals white, clawed, claw equaling calyx, limb deeply 2-fid into 2 spatulate lobes, to 5 mm, appendages ca. 0.5 mm;

stamens exserted, to 2 times length of calyx;

filaments usually dark purple;

stigmas 3, exserted, to 2 times length of calyx.

bisexual and unisexual, all flowers having bisexual flowers and staminate unisexual flowers, 6–8 mm diam.;

calyx prominently 10-veined, narrowly campanulate in flower, campanulate in fruit, 7–10(–11) × 4–6 mm, only slightly contracted around carpophore, papery and membranous, sparsely scabrid with short glandular-setose hairs, veins parallel, green, with pale commissures, margins dentate, lobes broadly ovate (as broad as long), 1–2 mm, not ciliate;

corolla cream or white, tinged purple, claw slightly exceeding calyx, narrowly triangular, auriculate, puberulent at base, limb oblong, 2-lobed, 2–4 mm, lobes entire, appendages 0.1–0.5 mm and broad, entire;

stamens equaling petals;

styles 3, equaling petals, reduced in functionally staminate flowers.

Capsules

ovoid, equaling calyx and sometimes splitting it, opening by 6 recurved, narrowly lanceolate teeth;

carpophore ca. 1 mm.

ovoid-ellipsoid, equaling calyx, opening by 6 spreading teeth;

carpophore ca. 2 mm.

Seeds

grayish brown, plump, broadly reniform, 0.6–1 mm, with concentric rings of papillae;

papillae slightly longer than broad.

brown, reniform-triangular, ca. 1 mm; papillate mainly around margin.

2n

= 24.

Silene csereii

Silene rectiramea

Phenology Flowering summer. Flowering April-May.
Habitat Cultivated fields, roadsides, waste land Pinyon pine woodland, dry slopes
Elevation 0-1600 m (0-5200 ft) 1800-2100 m (5900-6900 ft)
Distribution
from FNA
CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NY; OH; PA; SD; VT; WA; WI; WY; AB; BC; MB; ON; QC; SK; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Often confused with Silene vulgaris, S. csereii may be readily separated by the long, racemose primary branches of its inflorescence, the elliptic calyx that is constricted at both ends, tightly enclosing the capsule and lacking obvious reticulate venation, and the purple filaments.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Silene rectiramea is found in the Grand Canyon in Arizona. It is very similar to forms of S. verecunda and to S. thurberi. It is distinguished by its smaller flowers and fruiting calyces that are not or only slightly constricted around the carpophore and tend to be more sparsely pubescent with broader, shorter lobes. Also the appendages smaller and entire.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 180. FNA vol. 5, p. 200.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Name authority Baumgarten: Enum. Stirp. Transsilv. 3: 345. (1816) B. L. Robinson: Bot. Gaz. 28: 134. (1899)
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