Silene csereii |
Silene nivea |
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Balkan campion, Balkan catchfly, biennial campion, silène bisannuel |
evening campion, snowy campion |
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Habit | Plants annual or biennial, glabrous and somewhat glaucous; tap-root stout. | Plants perennial, rhizomatous; rhizome elongate. |
Stems | erect, sparingly branched below inflorescence, robust, to 65 cm. |
erect, simple to sparingly branched, leafy, 20–70 cm, glabrous to puberulent, especially distally. |
Leaves | basal few, usually withering by time of anthesis, blade spatulate; cauline numerous, 2 per node, blade 1-veined, ovate-lanceolate to oblanceolate, 3–7 cm × 7–30 mm, margins entire, apex acute. |
2 per node, sessile or short-petiolate, largest near mid-stem region, reduced and withering proximally, blade elliptic-lanceolate, base cuneate or rounded, apex gradually acuminate and acute, glabrous to puberulent. |
Inflorescences | many-flowered, open, bracteate; primary branches racemose, elongate, with sessile or shortly pedunculate cymes of 1–6 flowers per node; bracts narrowly lanceolate, 3–12 mm, hyaline-margined, apex acute. |
cymose, (1–)3–5(–12)-flowered, open, leafy. |
Pedicels | ascending, ± straight, 1–2 times calyx, broadening at calyx base. |
1/2–21/2 times calyx, apex often becoming deflexed, glabrous to hirsute. |
Flowers | calyx often obscurely ca. 20-veined, elliptic, abruptly contracted at base, opening constricted to 1/2 its diam., slightly inflated in flower, 7–10 × 3–4 mm, in fruit tightly enveloping capsule, ovoid, thin, enlarging to 9–13 × 5–7 mm, herbaceous, margins narrow, membranous, dentate with broadly triangular lobes to 1 mm, glabrous, veins obscure, usually purple tinged, without conspicuous, pale commissures, longitudinal, parallel, not obviously reticulate; petals white, clawed, claw equaling calyx, limb deeply 2-fid into 2 spatulate lobes, to 5 mm, appendages ca. 0.5 mm; stamens exserted, to 2 times length of calyx; filaments usually dark purple; stigmas 3, exserted, to 2 times length of calyx. |
calyx green, obscurely 10-veined, broadly tubular to campanulate, ± constricted at base around carpophore with broad umbilicate base, becoming broadly clavate in fruit, 14–17 × 5–9 mm, herbaceous, glabrous or hirsute, veins green, without pale commissures, lobes triangular, 2–3 mm; corolla white, clawed, claw equaling calyx, expanded distally into 2-lobed limb, limb oblong, 6–7 mm, appendages oblong, 1–1.6 mm, margins ± entire; stamens short-exserted; stigmas 3, short-exserted. |
Capsules | ovoid, equaling calyx and sometimes splitting it, opening by 6 recurved, narrowly lanceolate teeth; carpophore ca. 1 mm. |
globose, equaling calyx, opening by 3 broad teeth that sometimes split to form 6; carpophore 5–6 mm. |
Seeds | grayish brown, plump, broadly reniform, 0.6–1 mm, with concentric rings of papillae; papillae slightly longer than broad. |
dark brown to black, with grayish bloom, broadly reniform, not winged, 0.7–1 mm, sides with concentric crescents of low tubercles, larger and deeper on outer margins. |
2n | = 24. |
= 48. |
Silene csereii |
Silene nivea |
|
Phenology | Flowering summer. | Flowering late spring–summer. |
Habitat | Cultivated fields, roadsides, waste land | Alluvial woodlands |
Elevation | 0-1600 m (0-5200 ft) | 0-400 m (0-1300 ft) |
Distribution |
CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NY; OH; PA; SD; VT; WA; WI; WY; AB; BC; MB; ON; QC; SK; Europe [Introduced in North America]
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DC; IA; IL; IN; MA; MD; ME; MN; MO; OH; PA; VA; WI; WV
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Discussion | Often confused with Silene vulgaris, S. csereii may be readily separated by the long, racemose primary branches of its inflorescence, the elliptic calyx that is constricted at both ends, tightly enclosing the capsule and lacking obvious reticulate venation, and the purple filaments. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The green, obscurely veined, umbilicate calyx with its broad base constricted around the carpophore is unique among the North American members of the genus. Silene nivea is occasionally weedy. It was introduced near Québec City (ca. 1969) but did not persist, and probably is not native also in Maine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 180. | FNA vol. 5, p. 194. |
Parent taxa | Caryophyllaceae > subfam. Caryophylloideae > Silene | Caryophyllaceae > subfam. Caryophylloideae > Silene |
Sibling taxa | ||
Synonyms | Cucubalus niveus, S. alba | |
Name authority | Baumgarten: Enum. Stirp. Transsilv. 3: 345. (1816) | (Nuttall) Muhlenberg ex Otth: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 377. (1824) |
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