Silene campanulata |
Silene verecunda |
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bell catchfly, campanulate campion, red mountain catchfly, slender campion |
Dolores campion, San Francisco campion |
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Habit | Plants perennial; taproot stout; caudex much-branched, woody, producing many erect-to-straggling, little-branched flowering shoots. | Plants perennial; taproot stout; caudex branched, woody. | ||||
Stems | erect, 5–40 cm, softly pubescent to scabrous, eglandular or viscid-glandular, especially distally, very rarely glabrous, with several pairs of leaves equaling or shorter than internodes. |
usually several–many, rarely 1, erect, leafy, 10–55 cm, base often decumbent with marcescent leaf bases, scabrous-puberulent to pubescent, usually ± viscid-glandular distally, rarely densely so. |
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Leaves | sessile, or basal with short pseudopetiole; blade linear to lanceolate or broadly ovate, base round to cuneate, apex acute to shortly acuminate, puberulent on both surfaces, sometimes glandular. |
2 per node; basal petiolate, blade linear-lanceolate to lanceolate, 3–10 cm × 2–13 mm (including petiole), apex acute, glabrous to scabrous-puberulent or softly pubescent, petiole often ciliate; cauline sessile or petiolate, connate proximally, reduced distally, blade linear to narrowly lanceolate, 2–10 cm × 2–8 mm. |
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Inflorescences | usually with single dichotomy, rarely double, open, bracteate, branches often elongate, flowers 1 per node; bracts foliaceous. |
cymose with elongate ascending branches, open, flowers (1–)3 to many, bracteate, bracteolate; bracts and bracteoles lanceolate, 3–20 mm, apex acuminate. |
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Pedicels | sharply reflexed at base, especially after anthesis, equaling calyx. |
1/4–3 times longer than calyx, scabrous-puberulous to pubescent and glandular, ± viscid. |
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Flowers | nodding; calyx obscurely 10-veined, broadly campanulate, lobed, 6–8 mm, enlarging to 13–16 mm in fruit, herbaceous, usually with short, dense pubescence throughout, often glandular-viscid, veins green, rarely purplish tinged, conspicuous pale commissures absent; lobes ovate-triangular, 1/2 to equaling tube, herbaceous; petals creamy white, often greenish abaxially, rarely pink tinged to dusky pink (subsp. campanulata), clawed, to 2 times calyx, claw villose, limb deeply divided and fan-shaped with many narrow, linear lobes, lobes rapidly curling, margins deeply divided or erose, appendages 2–4, to 2 mm; stamens exserted; filaments hairy at base; styles 3, to 2 times calyx. |
calyx prominently 10-veined, tubular in flower, ± clavate and contracted around carpophore in fruit, 10–14 × 4–6 mm, margins dentate, veins parallel, green (rarely purplish), with pale commissures, lobes ascending, lanceolate, 2–3 mm, margins usually with obtuse, membranous border, apex spreading, shortly glandular-pubescent, usually viscid; corolla off-white (greenish) to dusky pink, clawed, claw equaling calyx, limb obovate, 2-lobed, 3–7 mm, shorter than claw, rarely with small lateral teeth, appendages 2, usually lacerate, 1–2 mm; stamens ca. equaling petals; styles 3(–4), often much longer than petals. |
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Capsules | ovoid, ca. equaling calyx and often splitting it, opening by 6 broadly triangular teeth; carpophore 1–2.5 mm. |
ovoid-ellipsoid, slightly longer than calyx, opening by 6 (or 8) recurved teeth; carpophore 2–5 mm. |
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Seeds | brown, reniform, 2–2.5 mm broad, coarsely and ± evenly papillate; papillae ca. as long as broad. |
dark reddish brown to black, reniform, ca. 1.5 mm, papillate, with larger papillae around margins. |
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2n | = 48. |
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Silene campanulata |
Silene verecunda |
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Phenology | Flowering summer. | |||||
Habitat | Meadows, chaparral, sagebush, open woodlands, dry pine forests, alpine ridges, dry canyons | |||||
Elevation | 0-3400 m (0-11200 ft) | |||||
Distribution |
CA; OR
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AZ; CA; NV; OR; UT; Mexico
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Silene verecunda is an exceptionally variable species, very difficult to circumscribe and tending to intergrade with S. bernardina, S. oregana, and S. grayi. It differs from the first two of those species mainly in having two-lobed petals. Silene grayi is a much smaller cespitose alpine plant with very large seeds. Hitchcock and Maguire divided S. verecunda into subsp. verecunda, subsp. platyota, and subsp. andersonii. Of these, subsp. andersonii is the most distinct, with a scabrous-puberulent indumentum, very narrow, stiff leaves, and rigid stems that are decumbent at the base, with marcescent leaf bases. The claw of the petals also is often more uniformly ciliate. Subspecies verecunda has a very different appearance, its mature calyx being shorter, broader, and markedly clavate. It is a short, stocky, viscid-glandular plant of exposed coastal habitats and may simply be a local ecotype. Subspecies platyota encompasses the remainder of the variation in the complex. Most of this variation consists of plants with fewer flowering stems; softer pubescence; broader, flat leaves; and thinner, more papery calyces. All these forms of S. verecunda appear to intergrade freely and, based on current information, any separation would be arbitrary. The species is in need of an experimental study to determine the nature of variation and its taxonomic value. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 176. | FNA vol. 5, p. 211. | ||||
Parent taxa | Caryophyllaceae > subfam. Caryophylloideae > Silene | Caryophyllaceae > subfam. Caryophylloideae > Silene | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | S. andersonii, S. behrii, S. luisana, S. occidentalis var. nancta, S. platyota, S. verecunda subsp. andersonii, S. verecunda var. eglandulosa, S. verecunda subsp. platyota, S. verecunda var. platyota | |||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 10: 341. (1875) | S. Watson: Proc. Amer. Acad. Arts 10: 344. (1875) | ||||
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