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bell catchfly, campanulate campion, red mountain catchfly, slender campion

Siberian catchfly

Habit Plants perennial; taproot stout; caudex much-branched, woody, producing many erect-to-straggling, little-branched flowering shoots. Plants perennial, producing several decumbent, short, woody, underground shoots bearing perennating buds and terminated by erect flowering shoots; taproot stout, woody; caudex crown branched.
Stems

erect, 5–40 cm, softly pubescent to scabrous, eglandular or viscid-glandular, especially distally, very rarely glabrous, with several pairs of leaves equaling or shorter than internodes.

simple but with short axillary leafy shoots, leafy, 40–60 cm, glossy, ± glabrous, with 10–20 nodes proximal to inflorescence.

Leaves

sessile, or basal with short pseudopetiole;

blade linear to lanceolate or broadly ovate, base round to cuneate, apex acute to shortly acuminate, puberulent on both surfaces, sometimes glandular.

basal withered at time of flowering;

cauline whorled, appearing fasciculate (more than 4) at each node, connate at base, reduced distally, tapered at both ends, blade linear to very narrowly elliptic-lanceolate, 0.5–8 cm × 1–12 mm, base broadened, margins scabrous, apex acute, subglabrous to minutely puberulent on both surfaces.

Inflorescences

usually with single dichotomy, rarely double, open, bracteate, branches often elongate, flowers 1 per node;

bracts foliaceous.

thyrsate, verticillate, open, with many short, ascending branches bearing large numbers of small, unisexual flowers, bracteate;

bracts connate basally, triangular, 2–10 mm, margins finely ciliate, apex acuminate.

Pedicels

sharply reflexed at base, especially after anthesis, equaling calyx.

ascending, straight, fasciculate, wiry, ± equaling calyx, glabrous or sparsely and minutely puberulent.

Flowers

nodding;

calyx obscurely 10-veined, broadly campanulate, lobed, 6–8 mm, enlarging to 13–16 mm in fruit, herbaceous, usually with short, dense pubescence throughout, often glandular-viscid, veins green, rarely purplish tinged, conspicuous pale commissures absent;

lobes ovate-triangular, 1/2 to equaling tube, herbaceous;

petals creamy white, often greenish abaxially, rarely pink tinged to dusky pink (subsp. campanulata), clawed, to 2 times calyx, claw villose, limb deeply divided and fan-shaped with many narrow, linear lobes, lobes rapidly curling, margins deeply divided or erose, appendages 2–4, to 2 mm;

stamens exserted;

filaments hairy at base;

styles 3, to 2 times calyx.

mostly unisexual;

calyx pale green, indistinctly 10-veined, elliptic, constricted near base, umbilicate, ca. 5 × 2 mm, lobes with thickened midrib, ovate-obtuse, 0.2 mm, margins crenate, membranous;

corolla greenish white, clawed, claw and limb not differentiated, narrowly oblong, unlobed, less than 2 times length of calyx, appendages absent;

stamens exserted, longer than corolla;

styles 3, exserted, longer than corolla.

Capsules

ovoid, ca. equaling calyx and often splitting it, opening by 6 broadly triangular teeth;

carpophore 1–2.5 mm.

green, ovoid-conic, longer than calyx, opening by 6 teeth;

carpophore shorter than 1 mm.

Seeds

brown, reniform, 2–2.5 mm broad, coarsely and ± evenly papillate;

papillae ca. as long as broad.

brown, winged, reniform, ca. 1 mm, faces with shallow radiating ridges.

Silene campanulata

Silene sibirica

Phenology Flowering summer.
Habitat Arable land, pastures, roadsides, sand dunes
Elevation 300-500 m (1000-1600 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
SK; Europe (Ukraine); Asia (Mongolia) [Introduced in North America]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene sibirica is a very distinct species with masses of small flowers on stiffly erect stems, and narrow, fascicled leaves. It was abundant and persisted for a number of years in the 1950s near Duck Lake and Bladworth in central Saskatchewan.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals dusky pink; leaf blades linear to lanceolate, usually less than 10 mm broad; stem pubescence scabrid, hairs 2 or 4 times as long as broad
subsp. campanulata
1. Petals creamy white; leaf blades lanceolate to ovate, usually more than 10 mm broad; stem pubescence not scabrid, hairs at least 5 times as long as broad
subsp. glandulosa
Source FNA vol. 5, p. 176. FNA vol. 5, p. 206.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. campanulata subsp. campanulata, S. campanulata subsp. glandulosa
Synonyms Cucubalus sibiricus
Name authority S. Watson: Proc. Amer. Acad. Arts 10: 341. (1875) (Linnaeus) Persoon: Syn. Pl. 1: 497. (1805)
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