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bell catchfly, campanulate campion, red mountain catchfly, slender campion

western campion, western catchfly

Habit Plants perennial; taproot stout; caudex much-branched, woody, producing many erect-to-straggling, little-branched flowering shoots. Plants perennial; taproot stout; caudex simple or branched, woody, bearing tufts of basal leaves.
Stems

erect, 5–40 cm, softly pubescent to scabrous, eglandular or viscid-glandular, especially distally, very rarely glabrous, with several pairs of leaves equaling or shorter than internodes.

erect, simple proximal to inflorescence, 30–60 cm, softly pubescent and stipitate-glandular.

Leaves

sessile, or basal with short pseudopetiole;

blade linear to lanceolate or broadly ovate, base round to cuneate, apex acute to shortly acuminate, puberulent on both surfaces, sometimes glandular.

2 per node;

basal ± petiolate, petiole ciliate, blade oblanceolate, spatulate, 5–12 cm × 7–20 mm, apex acute, short-pubescent on both surfaces;

cauline in 3(–4) pairs, reduced distally, blade oblanceolate to lanceolate.

Inflorescences

usually with single dichotomy, rarely double, open, bracteate, branches often elongate, flowers 1 per node;

bracts foliaceous.

open, narrow, with ascending branches, 9–25-flowered, bracteate, pubescent and stipitate-glandular;

bracts narrowly lanceolate, ciliate.

Pedicels

sharply reflexed at base, especially after anthesis, equaling calyx.

1/2–3 times longer than calyx.

Flowers

nodding;

calyx obscurely 10-veined, broadly campanulate, lobed, 6–8 mm, enlarging to 13–16 mm in fruit, herbaceous, usually with short, dense pubescence throughout, often glandular-viscid, veins green, rarely purplish tinged, conspicuous pale commissures absent;

lobes ovate-triangular, 1/2 to equaling tube, herbaceous;

petals creamy white, often greenish abaxially, rarely pink tinged to dusky pink (subsp. campanulata), clawed, to 2 times calyx, claw villose, limb deeply divided and fan-shaped with many narrow, linear lobes, lobes rapidly curling, margins deeply divided or erose, appendages 2–4, to 2 mm;

stamens exserted;

filaments hairy at base;

styles 3, to 2 times calyx.

calyx prominently 10-veined, tubular in flower and fruit, umbilicate, somewhat constricted around carpophore, 15–38 × 3–6 mm, papery, sparsely pubescent and stipitate-glandular, veins parallel, green, with pale commissures, lobes 5, broadly ovate, 2–4 mm, scarious around green midrib;

corolla pink or rose red (rarely white), clawed, claw equaling calyx, limb oblong, fanlike, deeply 4-lobed, lobes divergent, lanceolate (rarely with only 2 lobes, each with small lateral tooth), (7–)10–20 mm, appendages linear, 2–4 mm;

stamens exserted, shorter than petals;

stigmas 3, shorter than petals.

Capsules

ovoid, ca. equaling calyx and often splitting it, opening by 6 broadly triangular teeth;

carpophore 1–2.5 mm.

narrowly ovate-elliptic, longer than calyx, opening by 6 recurved teeth;

carpophore 4–18 mm.

Seeds

brown, reniform, 2–2.5 mm broad, coarsely and ± evenly papillate;

papillae ca. as long as broad.

grayish brown, reniform, 1–1.5 mm, verrucate.

2n

= 48.

Silene campanulata

Silene occidentalis

Phenology Flowering summer.
Habitat Grassy openings in chaparral, coniferous forests, and woodlands
Elevation 700-2300 m (2300-7500 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The long, tubular calyx and the proportionally long carpophore of Silene occidentalis are remarkable. Plants with the longest calyx tubes (more than 30 mm) have been referred to subsp. longistipitata and appear to be confined to Butte County. Calyx length varies greatly, however, and it is doubtful whether it is a justifiable basis for taxonomic recognition. The deeply lobed pink petals, together with its habit, give S. occidentalis a superficial resemblance to the European S. flos-cuculi, which occurs as an introduction on both sides of the North American continent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals dusky pink; leaf blades linear to lanceolate, usually less than 10 mm broad; stem pubescence scabrid, hairs 2 or 4 times as long as broad
subsp. campanulata
1. Petals creamy white; leaf blades lanceolate to ovate, usually more than 10 mm broad; stem pubescence not scabrid, hairs at least 5 times as long as broad
subsp. glandulosa
Source FNA vol. 5, p. 176. FNA vol. 5, p. 195.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. campanulata subsp. campanulata, S. campanulata subsp. glandulosa
Synonyms S. occidentalis subsp. longistipitata
Name authority S. Watson: Proc. Amer. Acad. Arts 10: 341. (1875) S. Watson: Proc. Amer. Acad. Arts 10: 343. (1875)
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