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bell catchfly, campanulate campion, red mountain catchfly, slender campion

Jan's catchfly, Nachlinger's catchfly, Nevada catchfly

Habit Plants perennial; taproot stout; caudex much-branched, woody, producing many erect-to-straggling, little-branched flowering shoots. Plants perennial; taproot stout; caudex usually branched, woody, producing tufts of basal leaves and 1–several flowering shoots.
Stems

erect, 5–40 cm, softly pubescent to scabrous, eglandular or viscid-glandular, especially distally, very rarely glabrous, with several pairs of leaves equaling or shorter than internodes.

subscapose, with (1–)2–3(–4) pairs of leaves, 6–25 cm, much reduced above base, retrorsely puberulent, eglandular.

Leaves

sessile, or basal with short pseudopetiole;

blade linear to lanceolate or broadly ovate, base round to cuneate, apex acute to shortly acuminate, puberulent on both surfaces, sometimes glandular.

2 per node;

basal blades oblanceolate, spatulate, 1.4–4 cm × 2–5 mm, base ciliate, apex acute, subglabrous to retrorse-puberulent;

cauline shortly connate proximally, blade narrowly oblanceolate to linear-lanceolate, 0.5–3.5 cm × 1–2 mm, apex purple tinged, retrorse-puberulent, often sparsely so.

Inflorescences

usually with single dichotomy, rarely double, open, bracteate, branches often elongate, flowers 1 per node;

bracts foliaceous.

1–3(–4)-flowered, open.

Pedicels

sharply reflexed at base, especially after anthesis, equaling calyx.

erect, 1–4 times longer than fruiting calyx, retrorsely puberulent, without glandular hairs.

Flowers

nodding;

calyx obscurely 10-veined, broadly campanulate, lobed, 6–8 mm, enlarging to 13–16 mm in fruit, herbaceous, usually with short, dense pubescence throughout, often glandular-viscid, veins green, rarely purplish tinged, conspicuous pale commissures absent;

lobes ovate-triangular, 1/2 to equaling tube, herbaceous;

petals creamy white, often greenish abaxially, rarely pink tinged to dusky pink (subsp. campanulata), clawed, to 2 times calyx, claw villose, limb deeply divided and fan-shaped with many narrow, linear lobes, lobes rapidly curling, margins deeply divided or erose, appendages 2–4, to 2 mm;

stamens exserted;

filaments hairy at base;

styles 3, to 2 times calyx.

calyx prominently 10-veined, cylindric, becoming narrowly ovoid, not inflated, not contracted proximally around carpophore, 6–11.5 × 3 mm in flower, broadening to ca. 5 mm in fruit, membranous, uniformly puberulent, without glandular hairs, veins parallel, green, with pale commissures, lobes triangular, 1–1.5 mm, margins narrowly membranous, apex purple-tipped;

corolla white, usually flushed with pink or purple towards apex, ligulate to ± oblanceolate, slightly longer than calyx, not clearly differentiated into claw and limb, unlobed to notched, auricles and appendages absent;

stamens equaling calyx;

styles 3, included in calyx.

Capsules

ovoid, ca. equaling calyx and often splitting it, opening by 6 broadly triangular teeth;

carpophore 1–2.5 mm.

ellipsoid-ovoid, slightly longer than calyx, opening by 6 ascending teeth;

carpophore ca. 1 mm.

Seeds

brown, reniform, 2–2.5 mm broad, coarsely and ± evenly papillate;

papillae ca. as long as broad.

rust colored, reniform, 0.7–1 mm, margins papillate;

papillae large, inflated.

Silene campanulata

Silene nachlingerae

Phenology Flowering summer.
Habitat Alpine limestone ridges and slopes
Elevation 2500-3000 m (8200-9800 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Silene nachlingerae is a small version of S. invisa, although the latter species has glabrous leaves. Both have the characteristic inflated (balloonlike) papillae around the outer edge of the seed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals dusky pink; leaf blades linear to lanceolate, usually less than 10 mm broad; stem pubescence scabrid, hairs 2 or 4 times as long as broad
subsp. campanulata
1. Petals creamy white; leaf blades lanceolate to ovate, usually more than 10 mm broad; stem pubescence not scabrid, hairs at least 5 times as long as broad
subsp. glandulosa
Source FNA vol. 5, p. 176. FNA vol. 5, p. 193.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. campanulata subsp. campanulata, S. campanulata subsp. glandulosa
Name authority S. Watson: Proc. Amer. Acad. Arts 10: 341. (1875) Tiehm: Brittonia 37: 344, fig. 1. (1985)
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