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bell catchfly, campanulate campion, red mountain catchfly, slender campion

Menzies' campion, Menzies' catchfly, Menzies' silene

Habit Plants perennial; taproot stout; caudex much-branched, woody, producing many erect-to-straggling, little-branched flowering shoots. Plants perennial, with several–many decumbent, sometimes cespitose or matted shoots; taproot slender.
Stems

erect, 5–40 cm, softly pubescent to scabrous, eglandular or viscid-glandular, especially distally, very rarely glabrous, with several pairs of leaves equaling or shorter than internodes.

decumbent to erect, simple or branched, leafy throughout, 5–30(–70) cm, usually glandular-puberulent distally, proximal pubescence varying from short and sparse to multicellular, crinkled and deflexed, glandular or not.

Leaves

sessile, or basal with short pseudopetiole;

blade linear to lanceolate or broadly ovate, base round to cuneate, apex acute to shortly acuminate, puberulent on both surfaces, sometimes glandular.

2 per node, sessile or short-petiolate;

blade usually oblanceolate to elliptic-lanceolate, rarely obovate or elliptic, 2–6(–10) cm × 3–20(–35) mm, broadest at or above middle, narrowed to base, margins ciliate with short, somewhat scabrid hairs, apex acute to acuminate, puberulent to pubescent.

Inflorescences

usually with single dichotomy, rarely double, open, bracteate, branches often elongate, flowers 1 per node;

bracts foliaceous.

cymose, or flowers axillary or solitary and terminal;

cyme loose, compound, leafy.

Pedicels

sharply reflexed at base, especially after anthesis, equaling calyx.

slender, 0.5–3 cm, glandular-puberulent.

Flowers

nodding;

calyx obscurely 10-veined, broadly campanulate, lobed, 6–8 mm, enlarging to 13–16 mm in fruit, herbaceous, usually with short, dense pubescence throughout, often glandular-viscid, veins green, rarely purplish tinged, conspicuous pale commissures absent;

lobes ovate-triangular, 1/2 to equaling tube, herbaceous;

petals creamy white, often greenish abaxially, rarely pink tinged to dusky pink (subsp. campanulata), clawed, to 2 times calyx, claw villose, limb deeply divided and fan-shaped with many narrow, linear lobes, lobes rapidly curling, margins deeply divided or erose, appendages 2–4, to 2 mm;

stamens exserted;

filaments hairy at base;

styles 3, to 2 times calyx.

functionally unisexual, usually bisexual;

calyx obscurely 10-veined, campanulate, 5–8 mm, ca. 1/2 as wide, herbaceous, margins dentate, lobes lanceolate, 1.5–3 mm, apex recurved, subacute to acuminate, puberulent to pilose and glandular, veins without conspicuous, pale commissures;

corolla white, clawed, 1–11/2 times calyx, claw shorter than calyx, limb oblong, 2-lobed, 1.5–3 mm, lobes oblong, apex obtuse, appendages 2, small, 0.1–0.4 mm;

stamens in functionally staminate flowers equaling corolla, otherwise reduced and included in calyx;

stigmas 3(–4), equaling corolla in functionally pistillate flowers, otherwise included in calyx, papillate along whole length.

Capsules

ovoid, ca. equaling calyx and often splitting it, opening by 6 broadly triangular teeth;

carpophore 1–2.5 mm.

green, becoming black, ovoid-ellipsoid, slightly longer than calyx, opening by 3 erect teeth which often split into 6;

carpophore ca. 1.5 mm.

Seeds

brown, reniform, 2–2.5 mm broad, coarsely and ± evenly papillate;

papillae ca. as long as broad.

black, not winged, broadly reniform, 0.5–1 mm, glossy, obscurely reticulate.

2n

= 24, 48.

Silene campanulata

Silene menziesii

Phenology Flowering throughout summer.
Habitat Common in open woodlands and forests, grasslands, gravelly places, river banks, mountains farther south
Elevation 200-3000 m (700-9800 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; MT; NE; NM; NV; OR; UT; WA; WY; AB; BC; MB; NT; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Silene menziesii is quite variable in the extent to which the inflorescence is developed and in its pubescence. This, coupled with the functionally dioecious nature of the species, has spawned a plethora of names, none of which appear to warrant recognition. The similar S. williamsii from Alaska and the Yukon Territory can be separated by its narrower lanceolate leaves that are broadest near the base and dull, usually brown, tuberculate seeds. Also, its stigmas are papillate only near the top. Silene seelyi is also very similar to S. menziesii but has dark red flowers and leaves that are smaller (to 2 cm in length) and broadest below the middle.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals dusky pink; leaf blades linear to lanceolate, usually less than 10 mm broad; stem pubescence scabrid, hairs 2 or 4 times as long as broad
subsp. campanulata
1. Petals creamy white; leaf blades lanceolate to ovate, usually more than 10 mm broad; stem pubescence not scabrid, hairs at least 5 times as long as broad
subsp. glandulosa
Source FNA vol. 5, p. 176. FNA vol. 5, p. 193.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. campanulata subsp. campanulata, S. campanulata subsp. glandulosa
Synonyms Anotites alsinoides, Anotites bakeri, Anotites costata, Anotites debilis, Anotites diffusa, Anotites discurrens, Anotites dorrii, Anotites elliptica, Anotites halophila, Anotites jonesii, Anotites latifolia, Anotites macilenta, Anotites menziesii, Anotites nodosa, Anotites picta, Anotites tenerrima, Anotites tereticaulis, Anotites villosula, Anotites viscosa, S. dorrii, S. menziesii subsp. dorrii, S. menziesii var. viscosa, S. obovata, S. stellarioides
Name authority S. Watson: Proc. Amer. Acad. Arts 10: 341. (1875) Hooker: Fl. Bor.-Amer. 1: 90, plate 30. (1830)
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