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silène muflier, sleepy campion, sleepy catch fly, sleepy catchfly campion, sleepy silene

Scouler's campion, Scouler's catchfly, Scouler's siliene, simple campion

Habit Plants annual; taproot slender. Plants perennial; taproot stout; caudex branched, woody, crowns 1–several.
Stems

erect, simple or branched, slender, to 80 cm, subglabrous to retrorsely puberulent especially proximally, distal internodes frequently glutinous.

erect, simple proximal to inflorescence, slender or stout, 10–80 cm, puberulent.

Leaves

2 per node;

blade with margins ciliate toward base, apex acute to obtuse;

basal blades oblanceolate, spatulate;

cauline narrowly oblanceolate to linear, 1–9 cm × 2–15 mm, scabrous or puberulent, rarely glabrous on both surfaces.

2 per node;

basal petiolate, blade oblanceolate, 6–25 cm × 4–30 mm, retrorsely puberulent on both surfaces;

cauline in 1–12 pairs, usually sessile, blade well developed, lanceolate to ovate-lanceolate, oblanceolate, or rarely linear or linear-lanceolate.

Inflorescences

cymose, open, branches usually ascending, several- to many-flowered, 1-flowered in depauperate specimens.

cymose, pseudo-racemose, or rarely paniculate, erect or nodding, with 1–12 flowering nodes, 2–20-flowered, open or dense, flowers paired or in many-flowered whorls, bracteate, cymes often sessile;

bracts 3–60 mm.

Pedicels

becoming deflexed at base of calyx, 1/4–2 times calyx, glandular-pubescent.

Flowers

mature calyx prominently 10-veined, campanulate to ovate, 5–9 × 3–5 mm, margins dentate, glabrous, veins parallel, with pale commissures;

lobes usually purple, triangular, acute, ca. 1 mm;

petals white, often suffused with dark red, rarely wholly dark red, limb ovate, usually 2-lobed, ca. 2.5 mm, slightly longer than calyx, rarely petals absent, claw narrow, appendages 0.1–0.4 mm;

stamens included;

styles 3;

stigmas included.

shortly pedicellate or sessile;

calyx prominently 10-veined, campanulate or tubular in flower, clavate, turbinate, or fusiform in fruit, constricted or not at base around carpophore in fruit, 8–20 × 3–8 mm, veins parallel, purplish or green, with pale commissures;

lobes lanceolate, 2–5 mm, apex obtuse with broad membranous margin and tip;

corolla white, greenish white, or pink, sometimes tinged pink or purple, clawed, claw longer than calyx, limb deeply 2–4-lobed, often with smaller lateral teeth, 2.5–8 mm, appendages 1–3 mm;

stamens ± equaling corolla claw;

styles 3–4, ± equaling corolla claw.

Capsules

equaling calyx, opening by 6 teeth;

carpophore less than 1 mm.

ovoid to ellipsoid, equaling or slightly longer than calyx, opening by 6 or 8 teeth;

carpophore 1.5–6 mm.

Seeds

dull gray-black, reniform, 0.5–0.8 mm diam., finely papillate.

brown or grayish brown, reniform, 1–1.5 mm, margins papillate, rugose on sides.

2n

= 24.

Silene antirrhina

Silene scouleri

Phenology Flowering spring–late summer.
Habitat Dry, sandy or gravelly places, roadsides, fields, waste places, open woods, often appearing after burning
Elevation 0-2300 m (0-7500 ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; ON; QC; SK; Mexico; South America; adventive in Europe
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

The six varieties and forms of Silene antirrhina noted above were named on the basis of stature and flower color, but none appear to be worthy of recognition. The species is very plastic, being greatly affected by moisture, exposure, and nutrients.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Silene scouleri is a very complex species that appears to be in the process of diverging into at least three different entities. Subspecies scouleri is a plant of the Pacific coast and lowlands. It has tall, stiffly erect stems, lanceolate to broadly lanceolate leaves, and a viscid inflorescence with many-flowered whorls of almost sessile flowers ranging in color from greenish white to rich pink. At the other extreme is subsp. pringlei, a plant of the mountains in Mexico extending northwards into Arizona and New Mexico. It has slender, somewhat nodding flowering stems with very narrow leaves. The flowers are usually paired at each node and secund on slender pedicels about equaling the calyx in length. The petals are off-white, sometimes tinged with dusky purple. Between the two extremes is subsp. hallii, a short, stocky plant of the Rocky Mountains and foothills with a few-flowered inflorescence. It has a larger, campanulate calyx, and some of the flowers usually become deflexed. Differentiation among these three forms is incomplete and plants indeterminate to subspecies are frequently encountered in areas away from the main distribution centers of the three subspecies. In northern Oregon and Idaho there appear to be populations connecting S. scouleri with S. oregana. They have some of the characteristics of S. oregana but not its laciniate petals. They may represent a more luxuriant form growing in taller vegetation, but their status needs further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyces campanulate, not or only slightly clavate in fruit, 13-18(-20) × (5-)6-8 mm; inflorescences with (1-)3-6(-8) flowering nodes; plants 10-40cm; pedicels stout
subsp. hallii
1. Calyces tubular to narrowly clavate in flower, clavate, turbinate, or fusiform in fruit, (8-)10-16 × 3.5-7 mm; inflorescences with 3-12 flowering nodes; plants 20-80 cm; pedicels slender
→ 2
2. Inflorescences erect, flowers in dense pseudowhorls of usually sessile, (2-)5-20-flowered cymes, both sessile and pedicellate flowers in each cyme; pedicels erect
subsp. scouleri
2. Inflorescences nodding, flowers usually paired at each node, all pedicellate; pedicels ± equaling calyx, often sharply deflexed at baseof calyx
subsp. pringlei
Source FNA vol. 5, p. 174. FNA vol. 5, p. 204.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. scouleri subsp. hallii, S. scouleri subsp. pringlei, S. scouleri subsp. scouleri
Synonyms S. antirrhina var. confinis, S. antirrhina var. depauperata, S. antirrhina var. divaricata, S. antirrhina var. laevigata, S. antirrhina var. subglaber, S. antirrhina var. vaccarifolia
Name authority Linnaeus: Sp. Pl. 1: 419. (1753) Hooker: Fl. Bor.-Amer. 1: 88. (1830)
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