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silène muflier, sleepy campion, sleepy catch fly, sleepy catchfly campion, sleepy silene

Drummond's campion, Drummond's catchfly, fork catchfly

Habit Plants annual; taproot slender. Plants perennial; taproot stout; caudex branched, somewhat fleshy.
Stems

erect, simple or branched, slender, to 80 cm, subglabrous to retrorsely puberulent especially proximally, distal internodes frequently glutinous.

erect, simple or several from base, retrorsely puberulent proximally, densely so and viscid distally, with stipitate glands.

Leaves

2 per node;

blade with margins ciliate toward base, apex acute to obtuse;

basal blades oblanceolate, spatulate;

cauline narrowly oblanceolate to linear, 1–9 cm × 2–15 mm, scabrous or puberulent, rarely glabrous on both surfaces.

blade with stiff, appressed pubescence on both surfaces;

basal petiolate, blade lanceolate to elliptic or oblanceolate, (1.5–)3–10 cm × 4–12 mm (including petiole);

cauline in 2–5 pairs, blade linear to linear-lanceolate, 3–9 cm × 2–7 mm.

Inflorescences

cymose, open, branches usually ascending, several- to many-flowered, 1-flowered in depauperate specimens.

1–20-flowered, bracteate, strongly viscid-glandular or less densely pubescent, longer hairs sometimes purple-septate;

bracts narrowly lanceolate, thick, 3–15 mm, herbaceous, apex acuminate.

Pedicels

stiffly erect, 0.1–5 cm, varying in length within same inflorescence.

Flowers

mature calyx prominently 10-veined, campanulate to ovate, 5–9 × 3–5 mm, margins dentate, glabrous, veins parallel, with pale commissures;

lobes usually purple, triangular, acute, ca. 1 mm;

petals white, often suffused with dark red, rarely wholly dark red, limb ovate, usually 2-lobed, ca. 2.5 mm, slightly longer than calyx, rarely petals absent, claw narrow, appendages 0.1–0.4 mm;

stamens included;

styles 3;

stigmas included.

calyx 10-veined, broadly tubular to narrowly ellipsoid, not inflated, 12–18 × 4–8 mm in fruit, 2–3 times as long as broad, membranous between veins, margins dentate with 5 triangular, 1.2–2 mm lobes erect in flower and spreading in fruit, apex acuminate, veins green;

petals off-white to dusky pink or dingy reddish purple, clawed, equaling or to 11/2 times calyx, claw broadened distally, limb not differentiated from claw, narrower than claw, 1–3 mm;

stamens included in calyx;

styles (4–)5, included in calyx.

Capsules

equaling calyx, opening by 6 teeth;

carpophore less than 1 mm.

12–15 mm, equaling calyx (rarely to 11/2 times calyx), opening by (4–)5 spreading teeth.

Seeds

dull gray-black, reniform, 0.5–0.8 mm diam., finely papillate.

dark brown, not winged, reniform to angular, 0.7–1 mm diam., margins finely papillate;

papillae triangular, slender, longer than broad.

2n

= 24.

Silene antirrhina

Silene drummondii

Phenology Flowering spring–late summer.
Habitat Dry, sandy or gravelly places, roadsides, fields, waste places, open woods, often appearing after burning
Elevation 0-2300 m (0-7500 ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; ON; QC; SK; Mexico; South America; adventive in Europe
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

The six varieties and forms of Silene antirrhina noted above were named on the basis of stature and flower color, but none appear to be worthy of recognition. The species is very plastic, being greatly affected by moisture, exposure, and nutrients.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Subspecies drummondii is characteristically a prairie taxon, while subsp. striata is associated with the Rocky Mountains. However, the two taxa frequently appear to intergrade; e.g., in the Cypress Hills of southern Alberta and Saskatchewan, and in the southern Rockies. Variety kruckebergii appears to be a luxuriant form with a more elongate capsule and calyx. Silene invisa, a Californian species, is similar to S. drummondii, some plants of which, from Nevada and Arizona, tend to be intermediate (see note under S. invisa).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals equaling calyx; fruiting calyces 12-15 × 4-6 mm, 21/ 4-3 times as long as broad; seeds ca. 0.7 mm diam.; inflorescences typically (1-)3-10(-20)-flowered
subsp. drummondii
1. Petals 11/ 1/ 2 times calyx and clearly exserted from it; fruiting calyces 13-18 × 6-8 mm, ca. 2 times as long as broad; seeds ca. 1 mm diam.; inflorescences typically 1-4(-8)-flowered
subsp. striata
Source FNA vol. 5, p. 174. FNA vol. 5, p. 183.
Parent taxa Caryophyllaceae > subfam. Caryophylloideae > Silene Caryophyllaceae > subfam. Caryophylloideae > Silene
Sibling taxa
S. acaulis, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. drummondii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
S. acaulis, S. antirrhina, S. aperta, S. armeria, S. bernardina, S. bridgesii, S. campanulata, S. caroliniana, S. chalcedonica, S. conica, S. coniflora, S. conoidea, S. coronaria, S. csereii, S. dichotoma, S. dioica, S. douglasii, S. flos-cuculi, S. gallica, S. grayi, S. hitchguirei, S. hookeri, S. invisa, S. involucrata, S. kingii, S. laciniata, S. latifolia, S. lemmonii, S. marmorensis, S. menziesii, S. nachlingerae, S. nivea, S. noctiflora, S. nuda, S. occidentalis, S. oregana, S. ostenfeldii, S. ovata, S. parishii, S. parryi, S. pendula, S. petersonii, S. plankii, S. polypetala, S. pseudatocion, S. rectiramea, S. regia, S. repens, S. rotundifolia, S. sargentii, S. scaposa, S. scouleri, S. seelyi, S. serpentinicola, S. sibirica, S. sorensenis, S. spaldingii, S. stellata, S. subciliata, S. suecica, S. suksdorfii, S. thurberi, S. uralensis, S. verecunda, S. virginica, S. viscaria, S. vulgaris, S. williamsii, S. wrightii
Subordinate taxa
S. drummondii subsp. drummondii, S. drummondii subsp. striata
Synonyms S. antirrhina var. confinis, S. antirrhina var. depauperata, S. antirrhina var. divaricata, S. antirrhina var. laevigata, S. antirrhina var. subglaber, S. antirrhina var. vaccarifolia Gastrolychnis drummondii, Lychnis drummondii, Melandrium drummondii, Wahlbergella drummondii
Name authority Linnaeus: Sp. Pl. 1: 419. (1753) Hooker: Fl. Bor.-Amer. 1: 89. (1830)
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