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three lobed oxytheca, three-lobed starry puncturebract, threelobe starry puncturebract

starry puncturebract

Habit Plants 0.7–5 × 0.7–6(–11) dm. Herbs, annual; taproots slender.
Stems

erect or more commonly spreading to prostrate.

arising directly from the root, spreading or prostrate, sometimes erect, solid, not fistulose or disarticulating into ringlike segments, glabrous or sparsely glandular.

Leaves

blades spatulate to oblanceolate or linear, 1–5(–9) × 0.2–0.7(–2) cm.

persistent or quickly deciduous, basal, rosulate;

petioles indistinct;

blade broadly linear or spatulate to oblanceolate, margins entire, strigose and glandular.

Inflorescences

open, 0.5–4 dm;

bracts mostly 2–8 × 1–3 mm, awns 0.8–1 mm.

terminal, cymose;

branches mostly dichotomous, not brittle or disarticulating into segments, round, glabrous or sparsely glandular;

bracts (2–)3(–4) at first node, 2–3 at distal nodes, distinct or connate, often positioned to side of node, scalelike, triangular or linear to ovate and 3-lobed, awned, sparsely glandular.

Peduncles

erect to spreading, 0.5–1.5 cm, glabrous except sparsely glandular proximally.

present or absent, erect to spreading.

Involucres

concolored, broadly turbinate, 3–8 mm, sparsely glandular;

teeth 5(–6), connate 1/3–1/2 their length;

awns greenish to reddish, 0.3–2 mm.

1 per node, not ribbed, tubular, narrowly turbinate to funnelform;

teeth 5(–6), awn-tipped.

Flowers

3–5(–10);

perianth white to pink or reddish, 2.5–4 mm;

tepals oblong, 3-lobed apically 1/3–1/2 their length, lobes sometimes laciniate;

filaments 1–4 mm;

anthers red to maroon, ellipsoid to oval, 0.5–0.7 mm.

2–5(–10) per involucre at any single time during full anthesis;

perianth white to rose or greenish yellow to red, funnelform when open, tubular when closed, hirsute and sparsely glandular abaxially;

tepals 6, connate 1/4–1/3 their length, monomorphic, 3–5-lobed or laciniate apically;

stamens 9;

filaments basally adnate, glabrous or minutely papillate basally;

anthers red to maroon, ellipsoid or oblong to oval.

Achenes

golden-brown, 1.2–2 mm.

usually included, golden- to red-brown, not winged, 3-gonous, glabrous.

Seeds

embryo curved.

x

= 20.

2n

= 40.

Sidotheca trilobata

Sidotheca

Phenology Flowering Apr–Sep.
Habitat Sandy flats, washes, slopes, chaparral communities, montane coniferous woodlands
Elevation 700-2100 m (2300-6900 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; nw Mexico
[BONAP county map]
Discussion

Sidotheca trilobata occurs in the Transverse Ranges of southern California from Los Angeles and San Bernardino counties southward into northern Baja California, Mexico. It is the most commonly encountered species of the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 3 (3 in the flora).

Sidotheca is allied to Eriogonum subg. Ganysma, approaching E. inerme in terms of foliar and overall habit. The trilobed to laciniate tepals resemble those of certain species of Chorizanthe. It is possible, as B. Ertter (1980) suggested, that the taxon was derived from Acanthoscyphus. In the 1950s, G. J. Goodman (1904–1999) proposed its recognition at generic rank, using the parahomonym “Neoxytheca.”

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres funnelform, white-margined, bracts connate more than 3/ 4 their length; tepals 3-5-lobed apically, lobes laciniate
S. emarginata
1. Involucres narrowly to broadly turbinate, concolored, bracts connate ca. 1/ 2 their length; tepals 3-lobed apically, lobes laciniate or not
→ 2
2. Perianths 1-2 mm, greenish yellow to red; tepals 3-lobed apically 1/ 5 their length; peduncles present or absent; awns of involucral bracts 0.3-1 mm; awns of inflorescence bracts 0.2-0.5 mm
S. caryophylloides
2. Perianths 2.5-4 mm, white to pink; tepals 3-lobed apically 1/ 1/ 2 their length; peduncles present; awns of involucral bracts 0.3-2 mm; awns of inflorescence bracts 0.8-1 mm
S. trilobata
Source FNA vol. 5, p. 440. FNA vol. 5, p. 439. Author: James L. Reveal.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Sidotheca Polygonaceae > subfam. Eriogonoideae
Sibling taxa
S. caryophylloides, S. emarginata
Subordinate taxa
S. caryophylloides, S. emarginata, S. trilobata
Synonyms Oxytheca trilobata, Eriogonum trilobatum Oxytheca section Neoxytheca
Name authority (A. Gray) Reveal: Harvard Pap. Bot. 9: 211. (2004) Reveal: Harvard Pap. Bot. 9: 211. (2004)
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