Sidalcea hirtipes |
Sidalcea pedata |
|
|---|---|---|
|
bluff mallow, bristly-stem checker-mallow, bristly-stem checkerbloom, hairy-stem checkermallow |
bird-foot checkerbloom |
|
| Habit | Herbs, perennial, usually in colonies, 0.7–1.3(–1.8) m, not glaucous, with thick, rather woody taproot and coarse, elongate (cordlike) rhizomes 20–100 × 5 mm. | Herbs, perennial, much of plant often tinted reddish purple, 0.2–0.4 m, not glaucous, with fleshy, simple to branched taproot, without rhizomelike rootstocks. |
| Stems | several, scattered, erect, arising from rhizome apices, usually slightly hollow, densely, harshly bristly-hirsute, hairs stiff, pustular, simple, forked, or stellate, often 2–2.5 mm. |
clustered, erect to slightly ascending, base erect to decumbent-ascending, usually unbranched, (plants nearly scapose), solid, long-bristly with hairs often 2 mm, sometimes also stellate-hairy near base, usually becoming finely hispid distally. |
| Leaves | basal and cauline; stipules linear-subulate, 6–8 × 1–1.5 mm; petioles of proximal leaves 20–30 cm, 3 times as long as blades, reduced distally to 1/2 times as long as blades; blades: basal and proximalmost orbiculate to reniform, shallowly 5–9-lobed, 10–15 × 10–15 cm, thick, base narrowly cordate, lobe margins coarsely crenate-dentate, apex rounded, surfaces coarsely hirsute, hairs stiff; distal orbiculate, deeply palmately 5–7-lobed, incised ± to base, lobes sometimes lobed again, base cuneate, apex acute, deeply 2–3-toothed, surfaces long-hirsute or with 2–4-rayed, stellate hairs abaxially. |
mostly basal, cauline 1–3; stipules lanceolate, 3–5 × 1 mm; petiole 6–9 cm, basal 2 times blade length, cauline 1/2 times blade length; blade usually orbiculate, 2–5(–6) × 2–5(–6) cm, base cordate, apex rounded, basal deeply dissected into 3–7 primary lobes, each less deeply incised and somewhat ternate, ultimate segments linear to oblong-elliptic, margins entire, distal blades repeatedly dissected with linear segments, surfaces densely hirtellous and stellate-hairy abaxially, less hairy with mostly simple hairs adaxially. |
| Inflorescences | erect, spiciform to subcapitate, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, proximals usually long-pedunculate, unbranched or branched, 20+-flowered, 10+ flowers usually open on spike at same time, not interrupted, not 1-sided, usually to 8 cm, usually not elongate but sometimes slightly elongated in some populations and to 20 cm in fruit; bracts paired or single, linear, distal undivided, proximal distinct to base, 6 mm, mostly slightly longer than pedicels. |
erect or ascending, spiciform, initially dense, later open, calyces not conspicuously overlapping except sometimes in fruit, usually unbranched, usually 20+-flowered, proximal flowers remotely, evenly spaced, elongated in fruit, often 1-sided if ascending and not 1-sided when strictly erect, 15–30(–40) cm, elongating with fruits spaced, axis often wine-red; bracts linear, usually undivided, 2.5–4 mm, usually ± equaling or longer than pedicels. |
| Pedicels | 1–3(–5) mm; involucellar bractlets absent. |
1–2(–3) mm; involucellar bractlets absent. |
| Flowers | bisexual or unisexual and pistillate, plants gynodioecious; calyx often purple tinted, 9–11 mm, to 11–16 mm in fruit, margins ciliate, hairs 1–2 mm, surfaces finely stellate-hairy at base and with coarser, longer, simple and stellate hairs apically; petals usually pale pink to rose-lavender, rarely white, slightly or not pale-veined, (9–)10–21 mm, pistillate often 9–14 mm; staminal column 5–7(–10) mm, hairy; anthers white; stigmas 5–10. |
bisexual or unisexual and pistillate, similar in size, plants gynodioecious; calyx usually wine-red, 4–5(–7) mm, usually not accrescent, finely stellate-puberulent, marginal hairs longer and often simple; petals dark rose-pink, sometimes pale-veined, 7–10(–12) mm; staminal column 3–5 mm, sparsely hairy; anthers white; stigmas 5–8. |
| Seeds | 2.5–3.5 mm. |
2 mm. |
| Schizocarps | 7–8 mm diam.; mericarps 5–10, 3.5–4 mm, glabrous or sparsely stellate-puberulent, roughened, prominently reticulate-veined, sides rugose and pitted, back less so, mucro 0.6–0.8 mm. |
5 mm diam.; mericarps 5–8, somewhat inflated, 2.5 mm, glabrous, smooth, back lightly grooved, not reticulate-veined or pitted, mucro 0.2–0.3 mm. |
| 2n | = 60. |
= 20. |
Sidalcea hirtipes |
Sidalcea pedata |
|
| Phenology | Flowering (Apr–)May–Jul(–Aug). | Flowering May–Aug. |
| Habitat | Prairie remnants, coastal bluffs, open shrublands, fencerows, meadows, usually mesic, basaltic soil | Moist meadows, open woodlands |
| Elevation | 0–200(–1200) m [0–700(–3900) ft] | (1500–)1600–2500 m [(4900–)5200–8200 ft] |
| Distribution |
OR; WA
|
CA |
| Discussion | Of conservation concern. Sidalcea hirtipes is uncommon and known from Clatsop, Lincoln, and Tillamook counties in Oregon and Clark, Lewis, and Wahkiakum counties in Washington. Its elevation and habitat vary, and it seems as much at home on steep coastal cliffs as in more inland, historic prairies and mountain meadows. Populations can appear to be large because of the long-rhizomatous and clonal nature of the plants; they are few and local. It is threatened by grazing, loss of habitat, fire suppression, road construction and maintenance, and changes in hydrology. It is a candidate for listing in Oregon and has been listed as endangered in Washington. Sidalcea hirtipes is characterized by its coarse indument of bristle hairs, its generally compact spikelike inflorescences, its relatively few, large, erect, hirsute leaves, and, especially, its extensive, coarse rhizomes. The inflorescences in some populations are elongated in fruit; its range, hirsute indument, and thick leaves along with coarse rhizomes help to distinguish it from other species. Stem internode length varies depending on habitat, as in many other Sidalcea. Molecular data suggest a relationship among S. hirtipes and S. asprella, S. celata, and S. gigantea (K. Andreasen and B. G. Baldwin 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea pedata is known from Bear Valley and Bluff Lake in San Bernardino County. It is generally easily distinguished by its relatively small flowers, its stature, its wine-red inflorescence axis, calyx, and buds, its fleshy taproot and lack of rhizomes, and its dense basal cluster of palmately dissected leaves. It is threatened by development, vehicles, and grazing, and has been listed as endangered in California and also federally. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Synonyms | S. spicata var. pedata | |
| Name authority | C. L. Hitchcock: Perenn. Sp. Sidalcea, 42. (1957) | A. Gray: Proc. Amer. Acad. Arts 22: 288. (1887) |
| Source | FNA vol. 6, p. 339. | FNA vol. 6, p. 351. |
| Web links | ||