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Sidalcea hirtipes

bluff mallow, bristly-stem checker-mallow, bristly-stem checkerbloom, hairy-stem checkermallow

California checkerbloom, checkerbloom, checkermallow, dwarf checker-mallow, dwarf checkerbloom, wild hollyhock

Habit Herbs, perennial, usually in colonies, 0.7–1.3(–1.8) m, not glaucous, with thick, rather woody taproot and coarse, elongate (cordlike) rhizomes 20–100 × 5 mm. Herbs, perennial, colonial or not, 0.2–0.6(–1.1) m, not glaucous, with woody caudex, usually with woody taproot, without rhizomes or rhizomes not cordlike, 3–10 mm diam.
Stems

several, scattered, erect, arising from rhizome apices, usually slightly hollow, densely, harshly bristly-hirsute, hairs stiff, pustular, simple, forked, or stellate, often 2–2.5 mm.

clustered or scattered, erect to ascending, or decumbent to suberect, sometimes rooting, unbranched or branched, solid, usually densely to sparsely hirsute, stellate-hairy, or glabrescent, hairs 1–2 mm, distal stem usually more sparsely hairy to glabrate.

Leaves

basal and cauline;

stipules linear-subulate, 6–8 × 1–1.5 mm;

petioles of proximal leaves 20–30 cm, 3 times as long as blades, reduced distally to 1/2 times as long as blades;

blades: basal and proximalmost orbiculate to reniform, shallowly 5–9-lobed, 10–15 × 10–15 cm, thick, base narrowly cordate, lobe margins coarsely crenate-dentate, apex rounded, surfaces coarsely hirsute, hairs stiff;

distal orbiculate, deeply palmately 5–7-lobed, incised ± to base, lobes sometimes lobed again, base cuneate, apex acute, deeply 2–3-toothed, surfaces long-hirsute or with 2–4-rayed, stellate hairs abaxially.

basal and/or cauline;

stipules green or purplish, linear-lanceolate or wide-lanceolate to oblong or ovate, 3–8(–12) × 1–3.5(–5) mm, often hairy to ciliate;

petioles of proximalmost leaves (5–)6–15(–30) cm, 3–9 times as long as blades, much reduced distally to 1/2–2 times as long as blade;

blades: proximalmost usually orbiculate, sometimes reniform, unlobed or shallowly lobed, (1–)4–15 × (1–)4–12 cm, base cordate, margins crenate, apex rounded, surfaces hairy, hairs stiff, either or both simple and stellate or forked, lobes cuneate-obovate, margins dentate;

mid stem unlobed or palmately 5–9-lobed, margins coarsely crenate-serrate, lobes toothed or lobed to dissected;

distalmost smaller, sometimes subsessile.

Inflorescences

erect, spiciform to subcapitate, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, proximals usually long-pedunculate, unbranched or branched, 20+-flowered, 10+ flowers usually open on spike at same time, not interrupted, not 1-sided, usually to 8 cm, usually not elongate but sometimes slightly elongated in some populations and to 20 cm in fruit;

bracts paired or single, linear, distal undivided, proximal distinct to base, 6 mm, mostly slightly longer than pedicels.

erect or somewhat ascending, dense, subcapitate or spicate especially when young to elongated and open, calyces overlapping or not, not long-pedunculate, usually unbranched, 2–21-flowered, usually light or bright pink to lavender or dark rose-purple, rarely white, pale-veined, 7–20(–30) mm, pistillate 7–11(–30) mm, bisexual 10–25(–30) mm;

staminal column 4–8 mm, hairy;

anthers white to pale pink or pale yellow;

stigmas (6 or)7 or 8(or 9).

Pedicels

1–3(–5) mm;

involucellar bractlets absent.

Flowers

bisexual or unisexual and pistillate, plants gynodioecious;

calyx often purple tinted, 9–11 mm, to 11–16 mm in fruit, margins ciliate, hairs 1–2 mm, surfaces finely stellate-hairy at base and with coarser, longer, simple and stellate hairs apically;

petals usually pale pink to rose-lavender, rarely white, slightly or not pale-veined, (9–)10–21 mm, pistillate often 9–14 mm;

staminal column 5–7(–10) mm, hairy;

anthers white;

stigmas 5–10.

Seeds

2.5–3.5 mm.

1.5–3 mm.

Schizocarps

7–8 mm diam.;

mericarps 5–10, 3.5–4 mm, glabrous or sparsely stellate-puberulent, roughened, prominently reticulate-veined, sides rugose and pitted, back less so, mucro 0.6–0.8 mm.

4–8 mm diam.;

mericarps (6 or)7 or 8(or 9), 2.5–4 mm, usually minutely hirtellous, stellate-puberulent, or glandular, rarely glabrous, roughened, sides ± honeycomb-pitted and reticulate-veined, back less so, mucro 0.3–1 mm.

2n

= 60.

= 20, 40, 60.

Sidalcea hirtipes

Sidalcea malviflora

Phenology Flowering (Apr–)May–Jul(–Aug).
Habitat Prairie remnants, coastal bluffs, open shrublands, fencerows, meadows, usually mesic, basaltic soil
Elevation 0–200(–1200) m [0–700(–3900) ft]
Distribution
from FNA
OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Sidalcea hirtipes is uncommon and known from Clatsop, Lincoln, and Tillamook counties in Oregon and Clark, Lewis, and Wahkiakum counties in Washington. Its elevation and habitat vary, and it seems as much at home on steep coastal cliffs as in more inland, historic prairies and mountain meadows. Populations can appear to be large because of the long-rhizomatous and clonal nature of the plants; they are few and local. It is threatened by grazing, loss of habitat, fire suppression, road construction and maintenance, and changes in hydrology. It is a candidate for listing in Oregon and has been listed as endangered in Washington. Sidalcea hirtipes is characterized by its coarse indument of bristle hairs, its generally compact spikelike inflorescences, its relatively few, large, erect, hirsute leaves, and, especially, its extensive, coarse rhizomes. The inflorescences in some populations are elongated in fruit; its range, hirsute indument, and thick leaves along with coarse rhizomes help to distinguish it from other species. Stem internode length varies depending on habitat, as in many other Sidalcea. Molecular data suggest a relationship among S. hirtipes and S. asprella, S. celata, and S. gigantea (K. Andreasen and B. G. Baldwin 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sidalcea malviflora includes local variants and intermediates. It was first described as a coastal taxon; later researchers subsequently included interior populations that shared a few characteristics. Molecular study has shown that the majority of the interior plants are not very closely related to the coastal forms; the species is again considered to be primarily coastal. Two inland taxa remain, subspp. californica and dolosa.

Sidalcea malviflora can generally be distinguished by its coastal distribution, its decumbent-based stems, its relatively short pedicels, its relatively large and showy petals that usually have conspicuous whitish veins, especially when dry, and its generally prominently reticulate-pitted and usually glandular-puberulent mericarps that have a mucro. The leaves can be extremely variable, from unlobed to highly dissected; the indument tends to be harsh to the touch.

Subspecies 7 (7 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Subordinate taxa
S. malviflora subsp. californica, S. malviflora subsp. dolosa, S. malviflora subsp. laciniata, S. malviflora subsp. malviflora, S. malviflora subsp. patula, S. malviflora subsp. purpurea, S. malviflora subsp. rostrata
Key
1. Inflorescences usually dense, flowers obviously overlapping, fruits either congested or spaced and not overlapping on elongated axis
→ 2
1. Inflorescences usually open, elongate, flowers not obviously overlapping except in bud, axis elongated in flower, fruit not congested
→ 3
2. Rhizomes freely rooting; leaves at midstem usually lobed; stems softly bristly-hairy proximally, stellate-hairy distally, usually not densely.
subsp. patula
2. Rhizomes not freely rooting; leaves at midstem unlobed; stems densely, softly bristly-hairy or stellate-hairy.
subsp. rostrata
3. Leaves: blades, except proximalmost, deeply lobed, lobes ternate or dissected, segments linear (Geranium-like); stem base usually decumbent, often rooting.
subsp. laciniata
5. Leaf blades: basal 1–2(–2.5) cm wide; calyces and stipules purplish; stems glabrous, short stellate-hairy, or sparsely bristly.
subsp. purpurea
5. Leaf blades: basal 2–6 cm wide; calyces and stipules usually green, not purple tinted (except sometimes at base of stipule only); stems softly bristly-hairy or stellate-hairy.
subsp. rostrata
6. Stem base hairy, hairs simple, soft, 2 mm; mericarps slightly reticulate-veined, wrinkled; rhizomes usually 10+ cm, freely rooting, sometimes matted; calyces sparsely stellate-puberulent and hirsute-bristly; San Bernardino Mountains.
subsp. dolosa
6. Stem base hairy, hairs stellate and/or simple, to 2 mm; mericarps strongly reticulate-veined, wrinkled or pitted; rhizomes usually to 10 cm, not freely rooting, not matted; calyces usually densely simple- or stellate-hairy; s outer South Coast and w Transverse ranges from Monterey to Los Angeles counties
→ 7
7. Stems usually stellate-hairy and/or spreading-bristly to glabrate; calyces densely stellate-puberulent and coarsely bristly; petals: bisexual 15–29 mm; coastal and insular.
subsp. malviflora
7. Stems densely, softly velvety stellate-hairy, hairs 1 mm; calyces densely stellate-puberulent; petals: bisexual 10–30 mm; inland.
subsp. californica
Synonyms Sida malviflora, Nuttallia malviflora
Name authority C. L. Hitchcock: Perenn. Sp. Sidalcea, 42. (1957) (de Candolle) A. Gray: Smithsonian Contr. Knowl. 3(5): 16. (1852)
Source FNA vol. 6, p. 339. Treatment author: Steven R. Hill. FNA vol. 6, p. 340. Treatment author: Steven R. Hill.
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