Sidalcea hickmanii |
Sidalcea celata |
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chaparral checkerbloom, Hickman's checkerbloom |
dwarf checkerbloom, Redding checkerbloom |
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Habit | Herbs, perennial, 0.1–0.8 m, not glaucous, with thick, woody taproot or caudex, without rhizomes. | Herbs, perennial, 0.4–0.8(–1) m, sometimes glaucous, with caudex and thick, woody rootstocks, not freely rooting, without rhizomes. | ||||||||||||||||||||
Stems | several to many (ca. 3–20+), clustered, erect to ascending, branched or unbranched, solid, usually densely stellate-canescent. |
mostly single, usually erect, sometimes branched distally, solid (sometimes hollow in older, robust plants), sometimes glaucous, proximally densely bristly-hairy, hairs reflexed, stiff, simple, sometimes also coarse, 2–3-rayed, stellate, 1–1.6(–2.5) mm, pustulate, sparser distally, with minute, stellate hairs in inflorescence. |
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Leaves | cauline, evenly arrayed on stem, usually similar in size, shape; stipules linear-lanceolate to ovate, 2–9 × 1–3 mm, widest above base, width sometimes exceeding stem diam.; petiole 0.6–3(–9) cm, 1/2–3 times as long as blade, apex often with pulvinus; blade orbiculate or reniform to flabelliform, unlobed and margins coarsely crenate to shallowly or deeply lobed, 1–7 × 1–7 cm, usually wider than long, base truncate or cordate, apex rounded, surfaces stellate-hairy. |
mostly basal, some cauline; stipules usually early-deciduous, linear-lanceolate, 3–7 × 1 mm; proximal petioles 15–18 cm, 3–4 times as long as blade, distal usually to 1/2 times to as long as blade; blades: basal rounded, usually palmately 7-lobed, shallowly incised, 4–8 × 4–8 cm, lobes 1.5–2 cm wide, margins crenate-dentate, 3–7-toothed, apices obtuse; distal 3–10, palmately 5-lobed, deeply incised, 2–6 × 2–6 cm, lobes narrow, linear, 2–3.5 × 2–4 mm, margins entire or 1–5-toothed, surfaces: abaxial stellate-hairy, hairs 6-rayed, 1.5 mm, adaxial hairy, hairs simple, 1.5 mm, distalmost glabrate adaxially and sometimes simple-hairy. |
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Inflorescences | erect, infrequently ascending, usually spiciform, dense or open, calyces overlapping or not, branched or unbranched, 2–20+-flowered, proximal flowers scattered, usually more congested distally, not notably elongate in flower, not 1-sided, (1.5–)3–25 cm, usually longer in fruit; bracts linear to ovate-lanceolate or oblong, undivided, 2-fid, or divided, 2–8(–12) mm, not involucrelike, distal entire to 2-fid, stipulelike, proximalmost not involucrelike, divided to base, much shorter than to nearly equaling calyx. |
erect, open, calyces not conspicuously overlapping except sometimes in bud, unbranched or branched, (5–)10–12(–23)-flowered, elongate, sometimes 1-sided, 10–40 cm; bracts lanceolate, usually undivided, 3–4 mm, shorter than pedicels and calyx. |
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Pedicels | 1–4(–5) mm; involucellar bractlets (2 or)3, 2–10 mm, shorter to slightly longer than calyx. |
(3–)5–10(–15) mm; involucellar bractlets absent. |
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Flowers | usually bisexual, infrequently unisexual and pistillate; calyx 4–12 mm, densely to sparsely stellate-puberulent to long-bristly; petals usually pale pink to pink-lavender, rarely white, veins not conspicuously whitened, 5–17 mm; staminal column 4–7 mm, hairy; anthers white to pale pinkish or pale yellow; stigmas (4–)6 or 7(–10). |
usually bisexual, infrequently unisexual and pistillate, plants gynodioecious; calyx (7–)9–10 mm, to 12–14 mm in fruit, densely stellate-puberulent; petals pale pinkish lavender, often pale-veined especially when dry, pistillate 10–20 mm, bisexual 20–25(–31) mm; staminal column 7–8(–10) mm, sparsely puberulent; anthers white; stigmas (6)7(8). |
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Seeds | 1–2 mm. |
2.5 mm. |
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Schizocarps | 4–7 mm diam.; mericarps usually (4–)6 or 7(–10), (1.5–)2–2.5 mm, glabrous, sides usually smooth, thin, margins and back usually lightly reticulate-veined, transversely corrugated, back usually with medial, raised line, not pitted, mucro absent. |
6–8 mm diam.; mericarps (6)7(8), 3–4 mm, minutely glandularpuberulent, glabrescent, margins sometimes sharpedged, roughened, strongly reticulate-veined, sides and back deeply pitted, honeycomblike, median line on back but not furrowed, mucro 1 mm. |
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2n | = 60. |
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Sidalcea hickmanii |
Sidalcea celata |
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Phenology | Flowering May–Jun(–Aug). | |||||||||||||||||||||
Habitat | Open oak woodlands, wet sites, sometimes on serpentine | |||||||||||||||||||||
Elevation | 100–400(–1600) m (300–1300(–5200) ft) | |||||||||||||||||||||
Distribution |
CA; OR
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CA |
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Discussion | Sidalcea hickmanii is found in isolated populations from southern California to southwestern Oregon and appears to have a relict distribution. K. Andreasen and B. G. Baldwin (2001, 2003) suggested that it is basal within Sidalcea. It is distinctive in having three (normally two in subsp. petraea) involucellar bractlets attached to the calyx, no mucro on the mericarps, and leaves that are almost the same size and shape throughout the stem. Each subspecies apparently represents a distinct relictual colony; the sexuality of these is not well known because of the paucity of specimens. As in many sidalceas, this species in particular appears to be fire-dependent. Subspecies 7 (7 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea celata has been recognized as a local species endemic to Shasta and Tehama counties (especially near Redding) in the northern inner North Coast Ranges; it was previously included within S. malviflora. Hitchcock’s concept of S. celata as a subspecies was broader and included plants from additional counties that do not match the type very well; most have been re-identified as S. asprella or S. oregana. Sidalcea celata has been confused with S. asprella, S. gigantea, and S. robusta and appears to be variable. Its narrow distribution, preference for serpentine, typical lack of elongated rhizomes, basally retrorsely hirsute stems, presence of basal leaves, and relatively large bisexual flowers and fruits are distinctive. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 334. | FNA vol. 6, p. 329. | ||||||||||||||||||||
Parent taxa | Malvaceae > subfam. Malvoideae > Sidalcea | Malvaceae > subfam. Malvoideae > Sidalcea | ||||||||||||||||||||
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Synonyms | S. malviflora var. celata, S. malviflora subsp. celata | |||||||||||||||||||||
Name authority | Greene: Pittonia 1: 139. (1887) | (Jepson) S. R. Hill: Madroño 56: 106. (2009) | ||||||||||||||||||||
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