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Hartweg's checkerbloom, valley checkerbloom

Henderson's checker-mallow, Henderson's checkerbloom

Habit Herbs, annual, 0.1–0.4(–0.6) m, not glaucous, with taproot. Herbs, perennial, 0.5–1.5 m, not glaucous, sparsely hairy to nearly glabrous throughout, with woody taproot and short, thick, ascending rootstock, usually also with compact rhizomes to 10 mm diam.
Stems

single, erect, unbranched or distally branched, often zigzag, solid, proximally usually glabrous or sparsely stellate-puberulent.

usually several, clustered, erect, usually purplish tinted especially near base, mostly unbranched, sometimes proximally hollow, base glabrous or sparsely hairy, hairs minute, simple or stellate, more densely scabrous distally.

Leaves

cauline;

stipules deciduous or inconspicuous, sometimes purplish, subulate, 1–1.5(–3) × 0.5–1 mm;

petiole 0.5–1.5(–3) cm, usually 1/2 times involucellar bractlets absent.

basal and cauline, ± fleshy;

stipules deciduous, often purplish, lanceolate to linear, 6–11 × 1–3 mm;

petiole often purplish tinted, those of basal leaves to 30 cm, proximalmost 3 times as long as blades, mid cauline 1–2 times as long as blades, distalmost to 1/2 times as long as blades and leaves subsessile;

basal blades orbiculate, unlobed or shallowly 5-lobed, 7–15 × 7–15 cm, margins crenate or dentate, ciliate, surfaces glabrous or sparsely hirsute, hairs sometimes forked, not stellate, mid cauline palmately 7–9-lobed, incised 1/2+ length, 7–12(–17) × 7–12(–17) cm, lobes cuneate, margins coarsely dentate to pinnatifid, apex toothed, surfaces: adaxial sparsely hairy or only on veins, ciliate, distal cauline leaves 3–5-parted nearly to base, lobes narrowly lanceolate to linear, margins coarsely dentate-laciniate.

Inflorescences

erect, spiciform, dense, especially when young, sometimes elongate, calyces usually conspicuously overlapping in flower and sometimes in fruit, usually branched, 20-flowered, usually several to many open at same time, not 1-sided, 5–20 cm;

bracts purplish, lanceolate to linear, undivided, rarely 2-fid, 4–5 mm, longer than pedicels, shorter than calyx.

Pedicels

1–4 mm, to 7–8 mm in fruit;

involucellar bractlets absent.

Flowers

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 8–10(–12) mm, not much enlarging in fruit, stellate-canescent, sparsely ciliate;

petals pink to rose-purple or white, pale-veined, often whitened at base, 18–20(–25) mm;

stamens: outer filaments incompletely connate, distally distinct, anthers not attached to connate portion of filaments;

staminal column 6–7 mm, hairy;

anthers white, stalked, aborted in pistillate flowers;

stigmas 6 or 7.

bisexual or often unisexual and pistillate, plants gynodioecious;

calyx somewhat urceolate, 8–9 mm, to 10–12 mm in fruit, lobes strongly reticulate-veined, sparsely stellate-hairy or proximally glabrous, lobe tips usually purple, rarely green, margins ciliate;

petals bright pinkish lavender, drying deep purple, veins not paler, bases not white, pistillate usually 8–10 mm, bisexual (10–)15–20 mm;

staminal column 6–7 mm, hairy;

anthers white;

stigmas (5–)7 or 8(or 9).

Seeds

1.5–2 mm.

3 mm.

Schizocarps

5–7 mm diam.;

mericarps 6 or 7, 2.5–4 mm, sides smooth, margins rugose, back reticulate-veined, deeply pitted especially on top, glabrous or glandular-puberulent, mucro 0.5–0.8(–1) mm.

8–9 mm diam.;

mericarps (5–)7 or 8(or 9), 4 mm, smooth, glabrous (rarely sparsely glandular-puberulent), thin-walled, apical margins sharp-edged, not winged, back sometimes slightly wrinkled, mucro 0.8–1.3 mm.

2n

= 20.

Sidalcea hartwegii

Sidalcea hendersonii

Phenology Flowering (Mar–)Apr–Jun. Flowering Jun–Aug.
Habitat Dry to moist, grassy hillsides, foothill woodlands, vernal pools, often on serpentine Coastal marshes, sandy to boggy tidal areas, upper beach meadows
Elevation 30–800(–1000) m (100–2600(–3300) ft) 0–50 m (0–200 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Sidalcea hartwegii is widespread in California from Tulare to Shasta counties, a range similar to those of the other vernal-pool annuals. It is most easily recognized by its stamen column, on which the anthers are borne on free portions of filaments, unlike in the other annual

species and more typical of the perennial species. This helps to support the suggestion that the annual species were separately derived and not part of a single lineage. Sidalcea hartwegii often grows with S. calycosa, from which it can be distinguished also by its rugose rather than deeply longitudinally furrowed dorsal mericarp surfaces.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sidalcea hendersonii is showy and distinctive; it has been confused with S. oregana subsp. spicata and with S. cusickii, to which it is similar and apparently closely related. Like S. cusickii, S. hendersonii generally has purplish tinted vegetative parts, compact inflorescences with flowers that are mostly open at the same time, and ciliate calyx lobes. Sidalcea hendersonii is generally considered vulnerable; the number of extant populations has declined in recent decades, especially in Oregon, where it is close to extirpation. It can be distinguished by its marshy seacoast habitat, its often hollow stems, its tendency to be glabrous, its dense, compound inflorescences, and its relatively large mericarps and mucro. It is the northernmost species of Sidalcea and is the one better adapted to brackish or saline marshes. Sidalcea hendersonii occurs from Douglas County, Oregon, to British Columbia, and a single specimen is known from Alaska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 333. FNA vol. 6, p. 334.
Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Synonyms S. hartwegii var. tenella, S. tenella
Name authority A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 20. (1849) S. Watson: Proc. Amer. Acad. Arts 23: 262. (1888)
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