Sidalcea campestris |
Sidalcea hendersonii |
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|---|---|---|
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meadow checker-mallow, meadow checkerbloom, meadow sidalcea |
Henderson's checker-mallow, Henderson's checkerbloom |
|
| Habit | Herbs, perennial, 0.5–2 m, sometimes glaucous in age, with thick, rather woody taproot and short rhizomes usually to 10 cm × 10 mm. | Herbs, perennial, 0.5–1.5 m, not glaucous, sparsely hairy to nearly glabrous throughout, with woody taproot and short, thick, ascending rootstock, usually also with compact rhizomes to 10 mm diam. |
| Stems | single or clustered, erect, base often decumbent-ascending, solid (proximally hollow on older stems), proximally densely bristly-hirsute, hairs simple or forked, 0.5–1 mm, sometimes mixed with minute, stellate hairs, sometimes glabrous and glaucous in age, hairs usually more appressed, simple, stellate, or sparse distally. |
usually several, clustered, erect, usually purplish tinted especially near base, mostly unbranched, sometimes proximally hollow, base glabrous or sparsely hairy, hairs minute, simple or stellate, more densely scabrous distally. |
| Leaves | mostly cauline; stipules lanceolate, 4–11 × 1–1.5 mm; petioles of proximal leaves 15–20 cm, 2–3 times as long as blades, distal reduced to 0.5–10 cm, 1/2 times to as long as blades; blade 10–15 × 10–15 cm, smaller distally, surfaces scabrid-hairy, hairs dense, simple or stellate, stiff, stellate hairs mostly on adaxial surfaces, proximal blades orbiculate, shallowly to deeply palmately 7–9-lobed, 5–15 × 5–15 cm, base cordate, margins coarsely crenate-serrate, lobes apically 2–5-toothed; distal cauline leaves variable, mid blades usually palmately divided nearly to base into 5–7 cuneate lobes, 15 × 15 cm, lobes deeply cut to laciniately dissected, distal blades divided into (3–)5–7 linear, marginally subentire segments, surfaces glabrescent or with few hairs on abaxial surface veins, ciliate. |
basal and cauline, ± fleshy; stipules deciduous, often purplish, lanceolate to linear, 6–11 × 1–3 mm; petiole often purplish tinted, those of basal leaves to 30 cm, proximalmost 3 times as long as blades, mid cauline 1–2 times as long as blades, distalmost to 1/2 times as long as blades and leaves subsessile; basal blades orbiculate, unlobed or shallowly 5-lobed, 7–15 × 7–15 cm, margins crenate or dentate, ciliate, surfaces glabrous or sparsely hirsute, hairs sometimes forked, not stellate, mid cauline palmately 7–9-lobed, incised 1/2+ length, 7–12(–17) × 7–12(–17) cm, lobes cuneate, margins coarsely dentate to pinnatifid, apex toothed, surfaces: adaxial sparsely hairy or only on veins, ciliate, distal cauline leaves 3–5-parted nearly to base, lobes narrowly lanceolate to linear, margins coarsely dentate-laciniate. |
| Inflorescences | erect, open, spiciform, calyces not conspicuously overlapping except sometimes in bud, often branched from distal leaves, 15+-flowered, elongate, not 1-sided, 10–35 cm, proximal flowers spaced several cm apart, not leafy-bracted; bracts linear, distal undivided or 2-fid, proximal separate to base, 4–10 mm, usually equaling or longer than pedicels. |
erect, spiciform, dense, especially when young, sometimes elongate, calyces usually conspicuously overlapping in flower and sometimes in fruit, usually branched, 20-flowered, usually several to many open at same time, not 1-sided, 5–20 cm; bracts purplish, lanceolate to linear, undivided, rarely 2-fid, 4–5 mm, longer than pedicels, shorter than calyx. |
| Pedicels | 3–6(–20) mm; involucellar bractlets absent. |
1–4 mm, to 7–8 mm in fruit; involucellar bractlets absent. |
| Flowers | bisexual or unisexual and pistillate, plants gynodioecious; calyx 5–9 mm, pistillate 5–7 mm, bisexual 6–9 mm, 8–10 mm in fruit, uniformly, densely stellate-hairy or with coarser, longer, stellate hairs to 2 mm; petals usually not overlapping, nearly white to pale pink or pale lavender, pistillate 9–12 mm, bisexual 13–25 mm; staminal column 5–7 mm, hairy; anthers white to pale pink; stigmas (6 or)7 or 8. |
bisexual or often unisexual and pistillate, plants gynodioecious; calyx somewhat urceolate, 8–9 mm, to 10–12 mm in fruit, lobes strongly reticulate-veined, sparsely stellate-hairy or proximally glabrous, lobe tips usually purple, rarely green, margins ciliate; petals bright pinkish lavender, drying deep purple, veins not paler, bases not white, pistillate usually 8–10 mm, bisexual (10–)15–20 mm; staminal column 6–7 mm, hairy; anthers white; stigmas (5–)7 or 8(or 9). |
| Seeds | 2.5 mm. |
3 mm. |
| Schizocarps | 7–8 mm diam.; mericarps (6 or)7 or 8, 3.5 mm, roughened, sides prominently reticulate-rugose and pitted, back less so and glandular-puberulent, mucro 0.5–1 mm. |
8–9 mm diam.; mericarps (5–)7 or 8(or 9), 4 mm, smooth, glabrous (rarely sparsely glandular-puberulent), thin-walled, apical margins sharp-edged, not winged, back sometimes slightly wrinkled, mucro 0.8–1.3 mm. |
| 2n | = 60. |
= 20. |
Sidalcea campestris |
Sidalcea hendersonii |
|
| Phenology | Flowering May–Aug. | Flowering Jun–Aug. |
| Habitat | Open shrublands, meadows, hedgerows, prairies | Coastal marshes, sandy to boggy tidal areas, upper beach meadows |
| Elevation | 40–200 m [130–700 ft] | 0–50 m [0–160 ft] |
| Distribution |
OR
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AK; OR; WA; BC
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| Discussion | Sidalcea campestris is one of the taller species of Sidalcea and can be distinguished also by its range, hirsute indument, long inflorescences with spaced, long-stalked flowers, narrow very pale petals, and deeply incised pinnatifid stem leaves. It has been confused with S. hirtipes, S. nelsoniana, and S. oregana; it differs especially in characters of rhizomes, inflorescences, and leaves. It is a candidate for listing as threatened or endangered in Oregon. Specimens from British Columbia and Washington identified as S. campestris are either S. campestris escaped from cultivation or S. hendersonii. It is known as a native only from the Willamette Valley area (Multnomah and Washington to Benton and Linn counties). Apparently, it was introduced near Seattle, Washington; it may not persist there. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea hendersonii is showy and distinctive; it has been confused with S. oregana subsp. spicata and with S. cusickii, to which it is similar and apparently closely related. Like S. cusickii, S. hendersonii generally has purplish tinted vegetative parts, compact inflorescences with flowers that are mostly open at the same time, and ciliate calyx lobes. Sidalcea hendersonii is generally considered vulnerable; the number of extant populations has declined in recent decades, especially in Oregon, where it is close to extirpation. It can be distinguished by its marshy seacoast habitat, its often hollow stems, its tendency to be glabrous, its dense, compound inflorescences, and its relatively large mericarps and mucro. It is the northernmost species of Sidalcea and is the one better adapted to brackish or saline marshes. Sidalcea hendersonii occurs from Douglas County, Oregon, to British Columbia, and a single specimen is known from Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Synonyms | S. asplenifolia, S. sylvestris | |
| Name authority | Greene: Bull. Calif. Acad. Sci. 1: 76. (1885) | S. Watson: Proc. Amer. Acad. Arts 23: 262. (1888) |
| Source | FNA vol. 6, p. 328. | FNA vol. 6, p. 334. |
| Web links | ||