Sidalcea campestris |
Sidalcea celata |
|
|---|---|---|
|
meadow checker-mallow, meadow checkerbloom, meadow sidalcea |
dwarf checkerbloom, Redding checkerbloom |
|
| Habit | Herbs, perennial, 0.5–2 m, sometimes glaucous in age, with thick, rather woody taproot and short rhizomes usually to 10 cm × 10 mm. | Herbs, perennial, 0.4–0.8(–1) m, sometimes glaucous, with caudex and thick, woody rootstocks, not freely rooting, without rhizomes. |
| Stems | single or clustered, erect, base often decumbent-ascending, solid (proximally hollow on older stems), proximally densely bristly-hirsute, hairs simple or forked, 0.5–1 mm, sometimes mixed with minute, stellate hairs, sometimes glabrous and glaucous in age, hairs usually more appressed, simple, stellate, or sparse distally. |
mostly single, usually erect, sometimes branched distally, solid (sometimes hollow in older, robust plants), sometimes glaucous, proximally densely bristly-hairy, hairs reflexed, stiff, simple, sometimes also coarse, 2–3-rayed, stellate, 1–1.6(–2.5) mm, pustulate, sparser distally, with minute, stellate hairs in inflorescence. |
| Leaves | mostly cauline; stipules lanceolate, 4–11 × 1–1.5 mm; petioles of proximal leaves 15–20 cm, 2–3 times as long as blades, distal reduced to 0.5–10 cm, 1/2 times to as long as blades; blade 10–15 × 10–15 cm, smaller distally, surfaces scabrid-hairy, hairs dense, simple or stellate, stiff, stellate hairs mostly on adaxial surfaces, proximal blades orbiculate, shallowly to deeply palmately 7–9-lobed, 5–15 × 5–15 cm, base cordate, margins coarsely crenate-serrate, lobes apically 2–5-toothed; distal cauline leaves variable, mid blades usually palmately divided nearly to base into 5–7 cuneate lobes, 15 × 15 cm, lobes deeply cut to laciniately dissected, distal blades divided into (3–)5–7 linear, marginally subentire segments, surfaces glabrescent or with few hairs on abaxial surface veins, ciliate. |
mostly basal, some cauline; stipules usually early-deciduous, linear-lanceolate, 3–7 × 1 mm; proximal petioles 15–18 cm, 3–4 times as long as blade, distal usually to 1/2 times to as long as blade; blades: basal rounded, usually palmately 7-lobed, shallowly incised, 4–8 × 4–8 cm, lobes 1.5–2 cm wide, margins crenate-dentate, 3–7-toothed, apices obtuse; distal 3–10, palmately 5-lobed, deeply incised, 2–6 × 2–6 cm, lobes narrow, linear, 2–3.5 × 2–4 mm, margins entire or 1–5-toothed, surfaces: abaxial stellate-hairy, hairs 6-rayed, 1.5 mm, adaxial hairy, hairs simple, 1.5 mm, distalmost glabrate adaxially and sometimes simple-hairy. |
| Inflorescences | erect, open, spiciform, calyces not conspicuously overlapping except sometimes in bud, often branched from distal leaves, 15+-flowered, elongate, not 1-sided, 10–35 cm, proximal flowers spaced several cm apart, not leafy-bracted; bracts linear, distal undivided or 2-fid, proximal separate to base, 4–10 mm, usually equaling or longer than pedicels. |
erect, open, calyces not conspicuously overlapping except sometimes in bud, unbranched or branched, (5–)10–12(–23)-flowered, elongate, sometimes 1-sided, 10–40 cm; bracts lanceolate, usually undivided, 3–4 mm, shorter than pedicels and calyx. |
| Pedicels | 3–6(–20) mm; involucellar bractlets absent. |
(3–)5–10(–15) mm; involucellar bractlets absent. |
| Flowers | bisexual or unisexual and pistillate, plants gynodioecious; calyx 5–9 mm, pistillate 5–7 mm, bisexual 6–9 mm, 8–10 mm in fruit, uniformly, densely stellate-hairy or with coarser, longer, stellate hairs to 2 mm; petals usually not overlapping, nearly white to pale pink or pale lavender, pistillate 9–12 mm, bisexual 13–25 mm; staminal column 5–7 mm, hairy; anthers white to pale pink; stigmas (6 or)7 or 8. |
usually bisexual, infrequently unisexual and pistillate, plants gynodioecious; calyx (7–)9–10 mm, to 12–14 mm in fruit, densely stellate-puberulent; petals pale pinkish lavender, often pale-veined especially when dry, pistillate 10–20 mm, bisexual 20–25(–31) mm; staminal column 7–8(–10) mm, sparsely puberulent; anthers white; stigmas (6)7(8). |
| Seeds | 2.5 mm. |
2.5 mm. |
| Schizocarps | 7–8 mm diam.; mericarps (6 or)7 or 8, 3.5 mm, roughened, sides prominently reticulate-rugose and pitted, back less so and glandular-puberulent, mucro 0.5–1 mm. |
6–8 mm diam.; mericarps (6)7(8), 3–4 mm, minutely glandularpuberulent, glabrescent, margins sometimes sharpedged, roughened, strongly reticulate-veined, sides and back deeply pitted, honeycomblike, median line on back but not furrowed, mucro 1 mm. |
| 2n | = 60. |
= 60. |
Sidalcea campestris |
Sidalcea celata |
|
| Phenology | Flowering May–Aug. | Flowering May–Jun(–Aug). |
| Habitat | Open shrublands, meadows, hedgerows, prairies | Open oak woodlands, wet sites, sometimes on serpentine |
| Elevation | 40–200 m [130–700 ft] | 100–400(–1600) m [300–1300(–5200) ft] |
| Distribution |
OR
|
CA |
| Discussion | Sidalcea campestris is one of the taller species of Sidalcea and can be distinguished also by its range, hirsute indument, long inflorescences with spaced, long-stalked flowers, narrow very pale petals, and deeply incised pinnatifid stem leaves. It has been confused with S. hirtipes, S. nelsoniana, and S. oregana; it differs especially in characters of rhizomes, inflorescences, and leaves. It is a candidate for listing as threatened or endangered in Oregon. Specimens from British Columbia and Washington identified as S. campestris are either S. campestris escaped from cultivation or S. hendersonii. It is known as a native only from the Willamette Valley area (Multnomah and Washington to Benton and Linn counties). Apparently, it was introduced near Seattle, Washington; it may not persist there. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea celata has been recognized as a local species endemic to Shasta and Tehama counties (especially near Redding) in the northern inner North Coast Ranges; it was previously included within S. malviflora. Hitchcock’s concept of S. celata as a subspecies was broader and included plants from additional counties that do not match the type very well; most have been re-identified as S. asprella or S. oregana. Sidalcea celata has been confused with S. asprella, S. gigantea, and S. robusta and appears to be variable. Its narrow distribution, preference for serpentine, typical lack of elongated rhizomes, basally retrorsely hirsute stems, presence of basal leaves, and relatively large bisexual flowers and fruits are distinctive. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Synonyms | S. asplenifolia, S. sylvestris | S. malviflora var. celata, S. malviflora subsp. celata |
| Name authority | Greene: Bull. Calif. Acad. Sci. 1: 76. (1885) | (Jepson) S. R. Hill: Madroño 56: 106. (2009) |
| Source | FNA vol. 6, p. 328. | FNA vol. 6, p. 329. |
| Web links | ||