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meadow checker-mallow, meadow checkerbloom, meadow sidalcea

annual checkerbloom, checker mallow, vernal pool checkerbloom

Habit Herbs, perennial, 0.5–2 m, sometimes glaucous in age, with thick, rather woody taproot and short rhizomes usually to 10 cm × 10 mm. Herbs, annual, rarely perennial, (0.2–)0.3–0.5(–0.9) m, not glaucous, with taproot or rhizomelike stolons to 35 blade; basal blades usually orbiculate, unlobed or slightly palmately lobed, 1–2(–10) × 2–5(–10) cm, wider than long, margins crenate, shiny; cauline blades rounded, deeply 5–9(–11)-lobed, 1–2 × 2–4 cm, lobes linear-elliptic to oblanceolate-obtuse, margins entire, ciliate, surfaces slightly hirsute.
Stems

single or clustered, erect, base often decumbent-ascending, solid (proximally hollow on older stems), proximally densely bristly-hirsute, hairs simple or forked, 0.5–1 mm, sometimes mixed with minute, stellate hairs, sometimes glabrous and glaucous in age, hairs usually more appressed, simple, stellate, or sparse distally.

Leaves

mostly cauline;

stipules lanceolate, 4–11 × 1–1.5 mm;

petioles of proximal leaves 15–20 cm, 2–3 times as long as blades, distal reduced to 0.5–10 cm, 1/2 times to as long as blades;

blade 10–15 × 10–15 cm, smaller distally, surfaces scabrid-hairy, hairs dense, simple or stellate, stiff, stellate hairs mostly on adaxial surfaces, proximal blades orbiculate, shallowly to deeply palmately 7–9-lobed, 5–15 × 5–15 cm, base cordate, margins coarsely crenate-serrate, lobes apically 2–5-toothed;

distal cauline leaves variable, mid blades usually palmately divided nearly to base into 5–7 cuneate lobes, 15 × 15 cm, lobes deeply cut to laciniately dissected, distal blades divided into (3–)5–7 linear, marginally subentire segments, surfaces glabrescent or with few hairs on abaxial surface veins, ciliate.

Inflorescences

erect, open, spiciform, calyces not conspicuously overlapping except sometimes in bud, often branched from distal leaves, 15+-flowered, elongate, not 1-sided, 10–35 cm, proximal flowers spaced several cm apart, not leafy-bracted;

bracts linear, distal undivided or 2-fid, proximal separate to base, 4–10 mm, usually equaling or longer than pedicels.

erect, spicate, dense, calyces sometimes overlapping, unbranched or branched, distal stem sometimes leafless, many-flowered, not 1-sided, 2–10 cm, elongating at maturity;

bracts green or purplish, ovate to wide-elliptic, usually 2-fid, sometimes undivided, 2–6(–12) × 2–5 mm.

Pedicels

3–6(–20) mm;

involucellar bractlets absent.

(2–)4–5 mm;

involucellar bractlets absent.

Flowers

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 5–9 mm, pistillate 5–7 mm, bisexual 6–9 mm, 8–10 mm in fruit, uniformly, densely stellate-hairy or with coarser, longer, stellate hairs to 2 mm;

petals usually not overlapping, nearly white to pale pink or pale lavender, pistillate 9–12 mm, bisexual 13–25 mm;

staminal column 5–7 mm, hairy;

anthers white to pale pink;

stigmas (6 or)7 or 8.

bisexual or infrequently unisexual and pistillate, plants gynodioecious;

calyx often purple tinted or scarious, 4–12 mm, silky strigose-bristly at base and on veins, stellate-puberulent;

petals usually pale purple or pink, rarely white, base pale or white, (9–)10–25 mm, pistillate shortest;

stamens: outer filaments connate to apex, tube funnel-like, with continuous rim to which unstalked anthers attach;

staminal column 4–9 mm, hairy in proximal 1/2;

anthers white to pale purple;

stigmas 5–9.

Seeds

2.5 mm.

2–3.5 mm.

Schizocarps

7–8 mm diam.;

mericarps (6 or)7 or 8, 3.5 mm, roughened, sides prominently reticulate-rugose and pitted, back less so and glandular-puberulent, mucro 0.5–1 mm.

5–9 mm diam.;

mericarps 5–9, often purple tinted, 2.5–4.5 mm, glabrous, not especially roughened, sides reticulate-veined, back deeply longitudinally grooved, mucro often appressed, 0.5 mm.

2n

= 60.

Sidalcea campestris

Sidalcea calycosa

Phenology Flowering May–Aug.
Habitat Open shrublands, meadows, hedgerows, prairies
Elevation 40–200 m [130–700 ft]
Distribution
from FNA
OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Sidalcea campestris is one of the taller species of Sidalcea and can be distinguished also by its range, hirsute indument, long inflorescences with spaced, long-stalked flowers, narrow very pale petals, and deeply incised pinnatifid stem leaves. It has been confused with S. hirtipes, S. nelsoniana, and S. oregana; it differs especially in characters of rhizomes, inflorescences, and leaves. It is a candidate for listing as threatened or endangered in Oregon. Specimens from British Columbia and Washington identified as S. campestris are either S. campestris escaped from cultivation or S. hendersonii. It is known as a native only from the Willamette Valley area (Multnomah and Washington to Benton and Linn counties). Apparently, it was introduced near Seattle, Washington; it may not persist there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sidalcea calycosa is variable and found locally in marshes and vernal pools in northern California. The annual, short-taprooted, subsp. calycosa varies in height and branching and is consistent in other vegetative and reproductive features. The perennial subsp. rhizomata has elongated, amphibious stolons somewhat like those of the fern Marsilea and is found in marshes near the coast. It and subsp. calycosa have the same fragile nature and similar morphology, and their distinctive fruits are essentially indistinguishable, having the only multifurrowed or striate surfaces in Sidalcea. Because S. calycosa is not at all fibrous and tough like the other perennials, it is keyed with the annuals.

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. asprella, S. calycosa, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Subordinate taxa
S. calycosa subsp. calycosa, S. calycosa subsp. rhizomata
Key
1. Plants annual, with taproot, stem base rarely decumbent, rooting; bracts 2–6 mm, not obscuring calyx, glabrous or ciliate; calyces 4–7 mm; petals (9–)10–25 mm; relatively widespread in n, c California.
subsp. calycosa
1. Plants perennial, with rhizomelike stolons rooting at nodes; bracts 8–12 mm, mostly obscuring calyx, silky-hairy; calyces 6–12 mm; petals 20–25 mm; restricted to n, c coastal California.
subsp. rhizomata
Synonyms S. asplenifolia, S. sylvestris
Name authority Greene: Bull. Calif. Acad. Sci. 1: 76. (1885) M. E. Jones: Amer. Naturalist 17: 875. (1883)
Source FNA vol. 6, p. 328. Treatment author: Steven R. Hill. FNA vol. 6, p. 326. Treatment author: Steven R. Hill.
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