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Sidalcea asprella

dwarf checkerbloom, harsh checker mallow, harsh checkerbloom

scadden flat checkerbloom

Habit Herbs, perennial, 0.1–1(–1.2) m, infrequently ± glaucous, with caudex or not, usually with freely-rooting fibrous rootstocks or rhizomes (5–)10–30 cm × 2–4 mm, matted or not. Herbs, perennial, 0.3–0.7 m, not glaucous, with slender, elongated rhizomes to 1 cm diam.
Stems

usually single, erect and sometimes supported by adjacent plants (sprawling), base prostrate or decumbent-ascending to erect, often rooting, solid, not brittle, sometimes ± glaucous distally, proximally stellate-hairy, glabrate, hairs minute or larger and coarse (never simple only), usually 4-rayed, 0.5–1 mm.

single or in dense patches, erect, solid, hairy, hairs simple, spreading, bristly.

Leaves

basal and/or cauline, similar in size and shape;

stipules linear to lanceolate, 2–3 × 1.1 mm;

petiole (1–)5–10(–15) cm, longest on proximal leaves, 1–4 times longer on proximal leaves to 1/2 times to as long as blade on distal leaves;

blade usually shallowly to deeply palmately 3–7-lobed usually halfway to base, proximal and distal cauline blades rounded to reniform, 2–3 × 2–5 cm, usually wider than long, base cordate to truncate, margins crenate, apex blunt or rounded, lobes narrowest at base, margins usually apically coarsely toothed, rarely entire, surfaces stellate-puberulent.

cauline, evenly arrayed on stem;

stipules asymmetric-ovate and auriculate, 10–20 × 10–20 mm, width exceeding stem diam.;

petioles of proximal leaves 7–10 cm, gradually reduced distally to 2 cm, 1–1 1/2 times blade length proximally to 1/2 blade length distally;

blade ovate to elliptic, unlobed, usually (4–)7–8 × (2.5–)4–5 cm, reduced distally, base cordate, margins crenate-serrate, apex acute to rounded, surfaces glabrous abaxially, sparsely hirsute adaxially.

Inflorescences

ascending or erect, often spiciform, open, calyces not overlapping in flower or fruit, unbranched or branched, 2–15(–30)-flowered, elongate in both flower and fruit, usually 1-sided, 6–11(–30) cm;

bracts leaflike to linear, usually 2-fid, (2–)3–5(–15) mm.

erect, capitate, ± dense, calyces overlapping, unbranched, 2–10-flowered, not elongate, not 1-sided, 3–5 cm;

proximalmost bracts involucrelike, similar to stipules in size and shape, narrowed to linear distally, usually divided to base, 10–20 mm, longer than pedicels, equaling or slightly shorter than calyx.

Pedicels

2–5(–10) mm;

involucellar bractlets absent.

1–2 mm;

involucellar bractlets 3, 8–12 mm, equaling or longer than calyx.

Flowers

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 5–12 mm, uniformly densely stellate-puberulent;

petals pink to pale purple, pale-veined, (5–)10–28 mm, pistillate flowers darker, 5–15 mm;

staminal column 4–5 mm, stellate-puberulent;

anthers white;

stigmas (6 or)7 or 8.

bisexual;

calyx 8 mm, sparsely to densely bristly, sometimes also stellate-hairy;

petals pink, usually pale-veined, 15 mm;

staminal column 4–6 mm, hairy;

anthers white;

stigmas 7 or 8.

Seeds

1.5–2.8 mm.

1.9 mm.

Schizocarps

6–8 mm diam.;

mericarps (6 or)7 or 8, 3–4 mm, usually glandular-puberulent to stellate-puberulent, sometimes glabrous, roughened, strongly reticulate-veined, sides and back pitted, mucro 0.5–1 mm.

4–5 mm diam.;

mericarps 7 or 8, 2 mm, glabrous, smooth, back with medial line, mucro absent.

Sidalcea asprella

Sidalcea stipularis

Phenology Flowering Jun–Aug.
Habitat Marshes
Elevation 700 m [2300 ft]
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Sidalcea asprella is variable and occurs from the central Sierra Nevada to southwestern Oregon. Typical plants in the central Sierra Nevada have weak, elongated stems that are often supported by neighboring vegetation; they lack simple recurved hairs at the stem base and may have either elongated rhizomes or a caudex. It has been confused with S. celata, S. elegans, S. gigantea, and S. glaucescens; formerly it was included within S. malviflora; molecular study has shown that it is different from S. malviflora. It belongs to a group including S. celata, S. elegans, S. gigantea, and S. hirtipes (K. Andreasen and B. G. Baldwin 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Sidalcea stipularis is the most distinctive and easily recognized species of Sidalcea; it is also among the rarest. It is listed as endangered in California. The ovate unlobed leaves are found in no other Sidalcea species. The relatively large stipules, bracts, and involucellar bractlets are also unique in Sidalcea, making the inflorescence appear involucrate. Studies by K. Andreasen and B. G. Baldwin (2001, 2003) suggested that it is one of the basal species within Sidalcea, and it has probably been long isolated. Sidalcea stipularis is known from the northern Sierra Nevada foothills in Nevada County.

Sidalcea stipularis is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants (0.3–)0.5–1(–1.2) m, with caudex or usually compact rootstocks or rhizomes to 10(–30) cm × 4 mm; leaves mostly cauline; inflorescences 8–15 (–30)-flowered, erect; stems erect, sometimes weak and supported by other vegetation, sometimes proximally decumbent.
subsp. asprella
1. Plants 0.1–0.3(–0.4) m, with rhizomes 5–20 cm × 2(–3) mm; leaves mostly basal; inflorescences usually 2–10(–19)-flowered, ascending; stems decumbent-ascending to erect, sometimes proximally prostrate.
subsp. nana
Source FNA vol. 6, p. 325. Treatment author: Steven R. Hill. FNA vol. 6, p. 355. Treatment author: Steven R. Hill.
Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. virgata
Subordinate taxa
S. asprella subsp. asprella, S. asprella subsp. nana
Synonyms S. malviflora subsp. asprella
Name authority Greene: Bull. Calif. Acad. Sci. 1: 78. (1885) J. T. Howell & G. H. True: Four Seasons 4(4): 20, fig. 16. (1974)
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