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Sidalcea asprella

dwarf checkerbloom, harsh checker mallow, harsh checkerbloom

Keck's checkerbloom

Habit Herbs, perennial, 0.1–1(–1.2) m, infrequently ± glaucous, with caudex or not, usually with freely-rooting fibrous rootstocks or rhizomes (5–)10–30 cm × 2–4 mm, matted or not. Herbs, annual, 0.1–0.4 m, not glaucous, with taproot.
Stems

usually single, erect and sometimes supported by adjacent plants (sprawling), base prostrate or decumbent-ascending to erect, often rooting, solid, not brittle, sometimes ± glaucous distally, proximally stellate-hairy, glabrate, hairs minute or larger and coarse (never simple only), usually 4-rayed, 0.5–1 mm.

single, erect, sometimes branched distally, solid, short-stellate-puberulent and long, soft bristly-hairy, distally with multicellular trichomes intermixed, hairs erect.

Leaves

basal and/or cauline, similar in size and shape;

stipules linear to lanceolate, 2–3 × 1.1 mm;

petiole (1–)5–10(–15) cm, longest on proximal leaves, 1–4 times longer on proximal leaves to 1/2 times to as long as blade on distal leaves;

blade usually shallowly to deeply palmately 3–7-lobed usually halfway to base, proximal and distal cauline blades rounded to reniform, 2–3 × 2–5 cm, usually wider than long, base cordate to truncate, margins crenate, apex blunt or rounded, lobes narrowest at base, margins usually apically coarsely toothed, rarely entire, surfaces stellate-puberulent.

cauline, 3–5 per stem;

stipules linear-filiform, undivided or rarely few-divided in robust plants, 3–5 × 1 mm;

petiole 2–4.5(–6) cm, usually 1–2 times as long as blade;

blades: proximalmost orbiculate, unlobed, 1–2.5 × 1–2.5 cm, base cordate with narrow sinus, margins crenate, apex rounded;

other proximals rounded, unlobed or shallowly palmately 7–9-lobed, 2.5–4.8(–6) × 2.5–4.8(–6) cm, margins coarsely crenate, sinus wide to narrow, surfaces stellate-hairy;

distals gradually reduced, orbiculate, more deeply 3-lobed, 1.5–2.5 × 1.5–2.5 cm, lobes obovate, apically widened to nearly truncate and narrowed to base, margins entire, apex 2–5-toothed, teeth subequal.

Inflorescences

ascending or erect, often spiciform, open, calyces not overlapping in flower or fruit, unbranched or branched, 2–15(–30)-flowered, elongate in both flower and fruit, usually 1-sided, 6–11(–30) cm;

bracts leaflike to linear, usually 2-fid, (2–)3–5(–15) mm.

erect to ascending, usually open, calyces not overlapping except possibly in bud, unbranched or infrequently branched, 5–12-flowered per branch, not especially elongate, usually 1-sided, 5–10 cm, to 5–15 cm in fruit, 1/2 times plant height;

bracts linear-filiform, undivided or rarely few-divided in robust plants, (3–)7–11 mm, longer than pedicels, usually much shorter than calyx, not involucrelike, densely stellate-hairy and pilose-hirsute.

Pedicels

2–5(–10) mm;

involucellar bractlets absent.

2–4 mm, to 5–6 mm in fruit;

involucellar bractlets absent.

Flowers

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 5–12 mm, uniformly densely stellate-puberulent;

petals pink to pale purple, pale-veined, (5–)10–28 mm, pistillate flowers darker, 5–15 mm;

staminal column 4–5 mm, stellate-puberulent;

anthers white;

stigmas (6 or)7 or 8.

bisexual;

calyx 8–10 mm, to 11–14 mm in fruit, lobe base within with conspicuous, purplish spot 1–2 mm wide, hairy, hairs glandular and non-glandular, multicellular as in inflorescence;

petals dark pink, without pale veins, with or without reddish basal spot, 10–22(–26) mm;

filaments connate to apex of tube;

staminal column 3–5 mm, hairy, hairs relatively long, simple;

anthers white;

stigmas 4 or 5.

Seeds

1.5–2.8 mm.

1.5–2 mm.

Schizocarps

6–8 mm diam.;

mericarps (6 or)7 or 8, 3–4 mm, usually glandular-puberulent to stellate-puberulent, sometimes glabrous, roughened, strongly reticulate-veined, sides and back pitted, mucro 0.5–1 mm.

5–7 mm diam.;

mericarps 4 or 5, usually tinted pink when fresh, 3–4 mm, usually glabrous, back reticulate-veined, pitted, with prominent midvein, mucro absent but with 1–5 minute bristles in its place.

Sidalcea asprella

Sidalcea keckii

Phenology Flowering Apr–May(–Jun).
Habitat Grassy slopes, tolerant of, not restricted to, serpentine
Elevation 70–700 m [230–2300 ft]
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Sidalcea asprella is variable and occurs from the central Sierra Nevada to southwestern Oregon. Typical plants in the central Sierra Nevada have weak, elongated stems that are often supported by neighboring vegetation; they lack simple recurved hairs at the stem base and may have either elongated rhizomes or a caudex. It has been confused with S. celata, S. elegans, S. gigantea, and S. glaucescens; formerly it was included within S. malviflora; molecular study has shown that it is different from S. malviflora. It belongs to a group including S. celata, S. elegans, S. gigantea, and S. hirtipes (K. Andreasen and B. G. Baldwin 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sidalcea keckii, federally listed as endangered, was once thought to have been extirpated (S. R. Hill 1993); it was rediscovered in 1992. It appears to have occurred historically in at least seven counties; extant populations are thought to be very few. It is found in the southern inner North Coast Ranges in Colusa, Napa, Solano, and Yolo counties, and in the Sierra Nevada foothills in Fresno, Merced, and Tulare counties. It is closely related and similar to S. diploscypha and is often mistaken for that species; differences between the two are discussed under 8. S. diploscypha. Some plants of S. keckii in Colusa, Solano, and Yolo counties have divided bracts like those of S. diploscypha. Sidalcea keckii is vulnerable to agricultural and residential land development.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants (0.3–)0.5–1(–1.2) m, with caudex or usually compact rootstocks or rhizomes to 10(–30) cm × 4 mm; leaves mostly cauline; inflorescences 8–15 (–30)-flowered, erect; stems erect, sometimes weak and supported by other vegetation, sometimes proximally decumbent.
subsp. asprella
1. Plants 0.1–0.3(–0.4) m, with rhizomes 5–20 cm × 2(–3) mm; leaves mostly basal; inflorescences usually 2–10(–19)-flowered, ascending; stems decumbent-ascending to erect, sometimes proximally prostrate.
subsp. nana
Source FNA vol. 6, p. 325. Treatment author: Steven R. Hill. FNA vol. 6, p. 339. Treatment author: Steven R. Hill.
Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Subordinate taxa
S. asprella subsp. asprella, S. asprella subsp. nana
Synonyms S. malviflora subsp. asprella
Name authority Greene: Bull. Calif. Acad. Sci. 1: 78. (1885) Wiggins: Contr. Dudley Herb. 3: 56, plate 13, figs. 2 – 6. (1940)
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