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Sidalcea asprella

dwarf checkerbloom, harsh checker mallow, harsh checkerbloom

giant checkerbloom

Habit Herbs, perennial, 0.1–1(–1.2) m, infrequently ± glaucous, with caudex or not, usually with freely-rooting fibrous rootstocks or rhizomes (5–)10–30 cm × 2–4 mm, matted or not. Herbs, perennial, (0.8–)2(–2.5) m, not glaucous except on stems, usually in colonies, with rhizomes to 40–60 cm × (6–)10 mm, glabrescent with reflexed-appressed bristle hairs 2.5 mm.
Stems

usually single, erect and sometimes supported by adjacent plants (sprawling), base prostrate or decumbent-ascending to erect, often rooting, solid, not brittle, sometimes ± glaucous distally, proximally stellate-hairy, glabrate, hairs minute or larger and coarse (never simple only), usually 4-rayed, 0.5–1 mm.

single, scattered, erect, usually purple tinted proximally, hollow especially towards base, pithy distally, 10–14 mm diam. just distal to base, often glaucous, proximally densely retrorsely bristly-hairy and stellate-hairy, hairs pustulate, 1.5–2.5 mm, distally sometimes glabrous.

Leaves

basal and/or cauline, similar in size and shape;

stipules linear to lanceolate, 2–3 × 1.1 mm;

petiole (1–)5–10(–15) cm, longest on proximal leaves, 1–4 times longer on proximal leaves to 1/2 times to as long as blade on distal leaves;

blade usually shallowly to deeply palmately 3–7-lobed usually halfway to base, proximal and distal cauline blades rounded to reniform, 2–3 × 2–5 cm, usually wider than long, base cordate to truncate, margins crenate, apex blunt or rounded, lobes narrowest at base, margins usually apically coarsely toothed, rarely entire, surfaces stellate-puberulent.

mostly cauline, basal usually absent;

stipules deciduous, with pink band on stem at base, narrowly lanceolate, (3.5–)5(–8) × 0.7(–1.5) mm;

petioles of proximal leaves 6–8 cm, those of midstem leaves 10–14 cm, 2 times to as long as blade, apex curved with swollen portion or pulvinus 5–6 × 1.8 mm;

blades of proximalmost cauline leaves rounded, shallowly, palmately 4- or 5-lobed, 0.5–0.7 × 0.5–0.7 cm, lobe margins irregularly, sparsely dentate, apices rounded, mid-cauline blades 5–7-lobed, 6.5–12 × 10–13 cm, wider than long, gradually reduced distally, lobes straplike, divided 3/4 to base, lobes 5.5 × 1.5–1.8 cm, margins coarsely dentate, surfaces sparsely, minutely hispid and stellate-puberulent, distal leaves deeply 5-lobed, otherwise similar in shape, leaves immediately below inflorescence greatly reduced, irregularly 2- or 3-lobed.

Inflorescences

ascending or erect, often spiciform, open, calyces not overlapping in flower or fruit, unbranched or branched, 2–15(–30)-flowered, elongate in both flower and fruit, usually 1-sided, 6–11(–30) cm;

bracts leaflike to linear, usually 2-fid, (2–)3–5(–15) mm.

erect, spiciform, open, calyces usually not overlapping, usually branched, branches 14–18 cm, each branch 10–20 flowered, not 1-sided, elongate, 5–20 cm;

bracts 1, usually undivided, narrowly lanceolate, often 2-fid, 2.5 mm, equaling or slightly longer than pedicel, much shorter than calyx.

Pedicels

2–5(–10) mm;

involucellar bractlets absent.

2–3(–5) mm;

involucellar bractlets absent.

Flowers

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 5–12 mm, uniformly densely stellate-puberulent;

petals pink to pale purple, pale-veined, (5–)10–28 mm, pistillate flowers darker, 5–15 mm;

staminal column 4–5 mm, stellate-puberulent;

anthers white;

stigmas (6 or)7 or 8.

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 5–6 mm, to 8 mm in fruit, uniformly, densely stellate-puberulent, surface obscured;

petals pale pink, pale-veined, pistillate 7–9 mm, bisexual (10–)14–20(–25) mm;

staminal column 6–8 mm, stellate-puberulent;

anthers white to cream or pale yellow;

stigmas (6)7 or 8.

Seeds

1.5–2.8 mm.

1.5 mm.

Schizocarps

6–8 mm diam.;

mericarps (6 or)7 or 8, 3–4 mm, usually glandular-puberulent to stellate-puberulent, sometimes glabrous, roughened, strongly reticulate-veined, sides and back pitted, mucro 0.5–1 mm.

6–7 mm diam.;

mericarps (6)7 or 8, sides 3 mm, thick, back and margins thick, rounded, reticulate-veined, pitted, back with prominent groove, top sparsely glandular-stellate-puberulent, mucro 1 mm.

Sidalcea asprella

Sidalcea gigantea

Phenology Flowering Jul–Sep.
Habitat Moist to wet, forested slopes, seeps, stream margins, meadows, coniferous forests
Elevation (600–)900–1700 m [(2000–)3000–5600 ft]
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Sidalcea asprella is variable and occurs from the central Sierra Nevada to southwestern Oregon. Typical plants in the central Sierra Nevada have weak, elongated stems that are often supported by neighboring vegetation; they lack simple recurved hairs at the stem base and may have either elongated rhizomes or a caudex. It has been confused with S. celata, S. elegans, S. gigantea, and S. glaucescens; formerly it was included within S. malviflora; molecular study has shown that it is different from S. malviflora. It belongs to a group including S. celata, S. elegans, S. gigantea, and S. hirtipes (K. Andreasen and B. G. Baldwin 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sidalcea gigantea is likely the tallest Sidalcea species; it can be distinguished also by its range (high Cascades and the northern high Sierra Nevada), retrorse hirsute stem bases, thick, hollow stems, and massive, hirsute rhizome systems. Most large patches appear to be clonal and generally produce either bisexual or unisexual, pistillate stems. Sidalcea gigantea is closely related to, and has been confused with, both S. asprella and S. celata. Its leaves are most like those of S. asprella in that they are usually similar in shape throughout the stem; the tall stems and massive rhizome systems distinguish it from S. asprella, as do its occurrence at higher elevations and its later flowering time.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants (0.3–)0.5–1(–1.2) m, with caudex or usually compact rootstocks or rhizomes to 10(–30) cm × 4 mm; leaves mostly cauline; inflorescences 8–15 (–30)-flowered, erect; stems erect, sometimes weak and supported by other vegetation, sometimes proximally decumbent.
subsp. asprella
1. Plants 0.1–0.3(–0.4) m, with rhizomes 5–20 cm × 2(–3) mm; leaves mostly basal; inflorescences usually 2–10(–19)-flowered, ascending; stems decumbent-ascending to erect, sometimes proximally prostrate.
subsp. nana
Source FNA vol. 6, p. 325. Treatment author: Steven R. Hill. FNA vol. 6, p. 332. Treatment author: Steven R. Hill.
Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Subordinate taxa
S. asprella subsp. asprella, S. asprella subsp. nana
Synonyms S. malviflora subsp. asprella
Name authority Greene: Bull. Calif. Acad. Sci. 1: 78. (1885) G. L. Clifton: Madroño 56: 285, figs. 1 – 3. (2010)
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