Sidalcea asprella |
Sidalcea cusickii |
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dwarf checkerbloom, harsh checker mallow, harsh checkerbloom |
Cusick's checkerbloom, Cusick's checkermallow, Cusick's sidalcea |
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Habit | Herbs, perennial, 0.1–1(–1.2) m, infrequently ± glaucous, with caudex or not, usually with freely-rooting fibrous rootstocks or rhizomes (5–)10–30 cm × 2–4 mm, matted or not. | Herbs, perennial, 0.4–1.8 m, not glaucous, with thick taproot and freely-rooting rhizomes usually 3–10 mm diam. | ||||
Stems | usually single, erect and sometimes supported by adjacent plants (sprawling), base prostrate or decumbent-ascending to erect, often rooting, solid, not brittle, sometimes ± glaucous distally, proximally stellate-hairy, glabrate, hairs minute or larger and coarse (never simple only), usually 4-rayed, 0.5–1 mm. |
usually several, clustered, erect, often purplish, often thick and hollow proximally, usually proximally ± scabrous, hairs stellate and distally stellate-hairy, sometimes glabrous. |
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Leaves | basal and/or cauline, similar in size and shape; stipules linear to lanceolate, 2–3 × 1.1 mm; petiole (1–)5–10(–15) cm, longest on proximal leaves, 1–4 times longer on proximal leaves to 1/2 times to as long as blade on distal leaves; blade usually shallowly to deeply palmately 3–7-lobed usually halfway to base, proximal and distal cauline blades rounded to reniform, 2–3 × 2–5 cm, usually wider than long, base cordate to truncate, margins crenate, apex blunt or rounded, lobes narrowest at base, margins usually apically coarsely toothed, rarely entire, surfaces stellate-puberulent. |
mostly cauline; stipules persistent or deciduous, lanceolate, 6–8 × 1–1.5 mm; proximal petioles 25–30 cm, 4–5 times as long as blade, reduced to 7–13 cm on midstem leaves and to 2–3 cm on distalmost leaves, these to 1/2 times as long as blades; basal blades wide-ovate to orbiculate, shallowly to deeply, palmately 5–7(–9)-lobed, 6–13 × 6–13 cm, lobe margins toothed, teeth rounded, sometimes Ribes-like, base cordate, apex rounded, surfaces: abaxial densely stellate-pubescent, adaxial glabrous or sparsely stellate-pubescent on veins; cauline blades round, divided nearly to base, 5–9-lobed, lobes again deeply incised to subentire, 10–20 × 10–20 cm, distalmost 5–7-cleft to base. |
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Inflorescences | ascending or erect, often spiciform, open, calyces not overlapping in flower or fruit, unbranched or branched, 2–15(–30)-flowered, elongate in both flower and fruit, usually 1-sided, 6–11(–30) cm; bracts leaflike to linear, usually 2-fid, (2–)3–5(–15) mm. |
erect, spiciform, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, usually 20–30-branched per primary stem, 20+-flowered, elongate, not 1-sided, most flowers usually open at same time, branches relatively short, each spike unit 6 cm, longer in fruit; bracts ovate-lanceolate to subulate, undivided, 1–6(–10) mm, subequal or longer than pedicels, much shorter than calyx. |
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Pedicels | 2–5(–10) mm; involucellar bractlets absent. |
1–2(–5) mm; involucellar bractlets absent. |
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Flowers | bisexual or unisexual and pistillate, plants gynodioecious; calyx 5–12 mm, uniformly densely stellate-puberulent; petals pink to pale purple, pale-veined, (5–)10–28 mm, pistillate flowers darker, 5–15 mm; staminal column 4–5 mm, stellate-puberulent; anthers white; stigmas (6 or)7 or 8. |
bisexual or unisexual and pistillate, pistillate plants more frequent than bisexual ones, some flowers may be staminate, plants gynodioecious; calyx urceolate with swollen base especially in young fruit, pistillate 6–8 mm, bisexual 6–10 mm, strongly reticulate-veined, usually finely stellate-puberulent, sometimes glabrate, lobes usually purple tinted; petals 7 or 8, pale pink, pinkish rose, or rose-purple, usually neither pale-veined nor white at base, pistillate 8–12(–14) mm, staminate or bisexual (10–)11–19(–23) mm; staminal column 5–6 mm, hairy; anthers white; stigmas 7 or 8. |
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Seeds | 1.5–2.8 mm. |
2 mm. |
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Schizocarps | 6–8 mm diam.; mericarps (6 or)7 or 8, 3–4 mm, usually glandular-puberulent to stellate-puberulent, sometimes glabrous, roughened, strongly reticulate-veined, sides and back pitted, mucro 0.5–1 mm. |
6 mm diam.; mericarps 7 or 8, 3 mm, apical margins ± rounded, sides smooth or slightly reticulate-veined, not pitted, back essentially smooth, very sparsely glandular-puberulent near tip, mucro 0.5–1 mm. |
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2n | = 20. |
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Sidalcea asprella |
Sidalcea cusickii |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Moist to wet, mostly black, adobe soil, lowland and mountain meadows, often with Juncus and Camassia | |||||
Elevation | 100–200(–500), 1000–1400 m [300–700(–1600), 3300–4600 ft] | |||||
Distribution |
CA; OR
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OR
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Discussion | Subspecies 2 (2 in the flora). Sidalcea asprella is variable and occurs from the central Sierra Nevada to southwestern Oregon. Typical plants in the central Sierra Nevada have weak, elongated stems that are often supported by neighboring vegetation; they lack simple recurved hairs at the stem base and may have either elongated rhizomes or a caudex. It has been confused with S. celata, S. elegans, S. gigantea, and S. glaucescens; formerly it was included within S. malviflora; molecular study has shown that it is different from S. malviflora. It belongs to a group including S. celata, S. elegans, S. gigantea, and S. hirtipes (K. Andreasen and B. G. Baldwin 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea cusickii is showy and distinctive. It has been confused with S. campestris, S. hendersonii, S. nelsoniana, S. oregana (subsp. spicata), S. setosa, and S. virgata. Molecular data support a close relationship with S. hendersonii and S. virgata (K. Andreasen and B. G. Baldwin 2003b). Hitchcock distinguished subsp. cusickii and subsp. purpurea, the latter with purple, ciliate calyx lobes and stem proximally glabrous and thought to be more closely related to S. hendersonii and S. nelsoniana. The two variations overlap in range and characteristics and are not recognized here. There appear to be two groups of populations at different elevation ranges with little overlap. Overall, S. cusickii can be distinguished by its range and by its spikelike, many-flowered, thick, compounded racemes in which essentially all of the flowers are open at the same time. It is also unusual in the preponderance of pistillate-flowered individuals in a given population and the presence of what appear to be truly staminate, rather than bisexual, flowers on some individuals. It is found in the Willamette and Umpqua valley region in Douglas, Josephine, and Lane counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 325. | FNA vol. 6, p. 330. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | S. malviflora subsp. asprella | S. cusickii subsp. purpurea, S. oregana var. cusickii | ||||
Name authority | Greene: Bull. Calif. Acad. Sci. 1: 78. (1885) | Piper: Proc. Biol. Soc. Wash. 29: 99. (1916) | ||||
Web links |