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comb seymeria, combleaf senna, Piedmont blacksenna

blacksenna

Habit Herbs, annual; hemiparasitic.
Stems

pubescent to villous, eglandular.

erect, not fleshy, glabrous or villous to scabrid.

Leaves

blade margins pinnatifid to 2-pinnatifid, pinnules lanceolate, surfaces not scabrid.

cauline, opposite;

petiole absent;

blade not fleshy, not leathery, margins entire, irregularly lobed, pinnatifid, or 2-pinnatifid.

Inflorescences

axillary, flowers 2 per axil;

bracts present.

Pedicels

3–5 mm.

present;

bracteoles absent.

Flowers

calyx lobes lanceolate;

corolla yellow, sometimes with maroon spots on adaxial lobes, externally pubescent, internally pubescent between lobes and in a ring at stamen insertion;

filaments tomentose to lanate distally, anthers dehiscing to 1/4 length.

sepals 5, calyx +/- radially symmetric, campanulate, lobes linear to lanceolate;

petals 5, corolla yellow often with red, maroon, or purple markings in throat, bilabiate, campanulate with spreading lobes, abaxial lobes 3, adaxial 2;

stamens 4, equal to subequal, filaments hairy proximally, glabrous or tomentose to lanate distally, anthers glabrous;

staminode 0;

ovary 2-locular, placentation axile;

stigma simple.

Capsules

symmetric, pyriform, glabrescent to densely tomentose.

dehiscence loculicidal.

Seeds

irregularly shaped, wings present.

30–40, brown, globular-ovoid or irregular, wings present or absent.

x

= 13.

2n

= 26.

Seymeria pectinata

Seymeria

Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC
[WildflowerSearch map]
[BONAP county map]
from USDA
Mexico; s United States; West Indies
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 17 (5 in the flora).

The phylogenetic position of Seymeria remains unclear. N. D. Young et al. (1999) placed Seymeria in an unresolved clade with Agalinis and Macranthera, among other genera. M. C. Neel and M. P. Cummings (2004) recovered a clade including Aureolaria, Brachystigma, Dasistoma, and Seymeria. A. D. Wolfe et al. (2005) placed Seymeria sister to Agalinis. J. R. Bennett and S. Mathews (2006) found that Seymeria is in a clade including Agalinis and Aureolaria, and also Esterhazya J. G. Mikan.

Seymeria is one of the few North American root-parasitic genera of economic importance in that S. cassioides is a parasite of pine species used for lumber in the southeastern United States (L. J. Musselman 1996).

The common name blacksenna is derived from the superficial resemblance to Senna (Fabaceae).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems pubescent to villous; capsules densely tomentose.
subsp. pectinata
1. Stems pubescent to puberulous; capsules glabrescent to pubescent.
subsp. peninsularis
1. Corollas externally pubescent or tomentose.
→ 2
2. Filaments tomentose to lanate distally, anthers dehiscing to 1/4 lengths; capsules densely tomentose or pubescent to glabrescent; seeds: wings present.
S. pectinata
2. Filaments glabrous distally, anthers dehiscing 1/2+ lengths; capsules pubescent; seeds: wings absent.
S. bipinnatisecta
1. Corollas externally glabrous.
→ 3
3. Leaves: blade margins 2-pinnatifid, pinnules filiform, surfaces not scabrid; capsules pyriform.
S. cassioides
3. Leaves: blade margins entire, 3-lobed, irregularly pinnatifid, or slightly 2-pinnatifid, pinnules linear to lanceolate, surfaces scabrid; capsules +/- falcate.
→ 4
4. Leaves: blade margins entire, 3-lobed, or irregularly pinnatifid, surfaces minutely scabrid; pedicels 6–10 mm; capsules glabrous.
S. falcata
4. Leaves: blade margins pinnatifid or slightly 2-pinnatifid, surfaces strongly scabrid; pedicels 1.5–4 mm; capsules glandular-pubescent.
S. scabra
Source FNA vol. 17, p. 564. FNA vol. 17, p. 562. Author: Christopher P. Randle.
Parent taxa Orobanchaceae > Seymeria Orobanchaceae
Sibling taxa
S. bipinnatisecta, S. cassioides, S. falcata, S. scabra
Subordinate taxa
S. pectinata subsp. pectinata, S. pectinata subsp. peninsularis
S. bipinnatisecta, S. cassioides, S. falcata, S. pectinata, S. scabra
Synonyms Afzelia pectinata Afzelia
Name authority Pursh: Fl. Amer. Sept. 2: 737. (1813) Pursh: Fl. Amer. Sept. 2: 736. (1813) — name conserved
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