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wand riverhemp

poisonbean, rattlebush

Habit Shrubs or trees, to 4 m. Shrubs or subshrubs, to 3 m.
Stems

glabrous or pilose, hairs persistent, close-pressed, golden or clear in age, developing leaves and young stems with same pubescence of simple hairs;

pith solid or spongy.

glabrous in age;

pith spongy.

Leaves

13–25+ cm;

stipules 0.3–0.4 cm, with conspicuous inner fold throughout, hairs dense, close-pressed, inner fold and base with stipitate, multicellular glands;

pulvinus slightly more than 1/2 as long as petiole;

rachis ± sericeous, without stipitate glands, obscure gland(s) present at petiolule base;

stipels reduced in size between successive leaflets, long-filamentous glandular;

leaflets 28–36+, blades elliptic-ovate to oblong, base acute, apex obtuse to acute, surfaces silky sericeous abaxially, usually glabrous adaxially.

12–19 cm;

stipules 0.2–0.4 cm, with inconspicuous inner fold throughout, inner fold and base with stipitate, multicellular glands or glabrous;

pulvinus less than 1/2 as long as petiole;

rachis ± glabrous, without stipitate glands in canal;

stipels narrow obscure glands;

leaflets 16–50+, blades elliptic-ovate to oblong, base acute, apex obtuse, surfaces glaucous, usually glabrous abaxially, sometimes with diffuse, close-pressed hairs.

Inflorescences

5–15+-flowered, racemes.

1–12-flowered, racemes.

Peduncles

(0.6–)1.7–2.1(–4.1) cm.

(0.9–)1.7–1.8(–3) cm.

Flowers

(0.7–)0.9–1(–1.3) cm;

calyx ± zygomorphic, lobes 5, short-acuminate, rim of tube villose, stalked glands between abaxial lobes absent in fruit;

corolla yellow, banner venation sometimes darker;

banner ovate, base cordate-truncate, apex emarginate, becoming strongly reflexed and contorted, calluses as relatively small, acute teeth at claw base, thickened, knoblike at base of blade/top of claw;

wings without basal tooth;

keel ± same color throughout, apex acute, curved strongly inward, without basal tooth;

stamens incurved within keel;

style recurved;

ovules 4–6.

(0.9–)1.3–1.5(–1.7) cm;

calyx ± zygo-morphic, deciduous before fruits mature, lobes 5, short-deltate, rim of tube with spreading hairs, stalked glands usually present between abaxial lobes;

corolla pale yellow;

banner ovate, base truncate-obcordate, apex emarginate, calluses as relatively small ridges at claw base, forming a pocket, thickened, knoblike at top of claw;

wings without basal tooth;

keel ± same color throughout, apex rounded-obtuse, curved upward to slightly outward in apical tooth, without basal tooth;

stamens curved upward;

style curved upward with stamens;

ovules 4–9.

Legumes

red- to gray-brown, without horizontal mottling in age, 4-angled, square in cross section, straight or slightly falcate, (0.8–)4.4–5.5(–6.5) × (0.7–)0.8(–0.9) cm, thick, woody, seed chambers apparent in young fruits becoming obscure at maturity, margins of young fruits with shallow thin ridges resembling early wings of S. punicea or S. drummondii, ridges becoming thickened and rounded at maturity, beak short-pyramidal, (0.2–)0.4–0.7(–1.3) cm, indehiscent;

stipe (0.4–)0.5–0.6(–0.9) cm.

light tan to reddish brown, 4-angled, with pronounced torulose wings, square in cross section, straight, (3.2–)5.7–6.2(–9.3) × 0.8(–0.9) mm, seed chambers clearly visible externally, beak short- to long-pyramidal, (0.4–)0.7–0.8(–1.3) cm, indehiscent or tardily dehiscent;

stipe (1.1–)1.6(–2) cm.

Seeds

(1–)4 or 5(or 6), reddish brown to gray, without mottling, reniform-orbicular.

(1–)3 or 4(–9), dark reddish brown, without mottling, reniform-orbicular.

2n

= 12.

= 12.

Sesbania virgata

Sesbania drummondii

Phenology Flowering early summer–early fall. Flowering summer–fall.
Habitat Wet areas, ripar­ian, wetlands, coastal, disturbed sites. Wet areas, riparian on sandy soils, coastal sites, disturbed sites.
Elevation 0–20 m. [0–70 ft.] 0–600 m. [0–2000 ft.]
Distribution
from FNA
FL; MS; South America [Introduced also in se Mexico (Veracruz), West Indies, Central America]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MS; SC; TX; Mexico (Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Sesbania virgata is native to northern Argentina and southern Paraguay and has been introduced to numerous port cities in the Americas. In the flora area, it is known from the Pensacola, Florida, region and from several populations along the coastline southeastward to Hillsborough and Pinellas counties and westward to Harrison County, Mississippi. The most distinguishing characteristic of the species is the quadrangular pod.

Sesbania affinis De Wildeman (1904) is a later homonym (not Schrader ex de Candolle 1825) that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sesbania drummondii is often mistaken for the Mexican species S. cavanillesii S. Watson (= Sesbania longifolia de Candolle). It is distinguished by rounded versus acute leaf apices and winged versus not winged legumes. In bloom, the yellow-orange corollas make S. drummondii clearly distinct from S. punicea; in fruit, identification is simplified by the acuminate versus pyramidal beak and wavy versus straight-edged wings.

Daubentonia texana Pierce is a superfluous name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Sesbania Fabaceae > subfam. Faboideae > Sesbania
Sibling taxa
S. drummondii, S. grandiflora, S. herbacea, S. punicea, S. sericea, S. vesicaria
S. grandiflora, S. herbacea, S. punicea, S. sericea, S. vesicaria, S. virgata
Synonyms Aeschynomene virgata, Agati virgata, Coursetia virgata, Emerus marginatus, S. marginata, S. tetragona Daubentonia drummondii
Name authority (Cavanilles) Poiret in J. Lamarck et al.: Encycl. 7: 129. (1806) — (as Sesban) (Rydberg) Cory: Rhodora 38: 406. (1936)
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