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Roemer senna, twoleaf senna, twoleaf wild sensitive plant

blunt leafed senna, Chinese senna, Java-bean, sickle-pod wild senna, sicklepod

Habit Herbs, perennial, to 0.7 m. Leaves slightly sclerophyllous, 2.5–9.5 cm, hairy; stipules caducous; extrafloral nectary 1, between leaflet pair, shortly stipitate; leaflet pairs 1, blades lanceolate-oblong or lanceolate, 20–70 × 4–14 mm. Herbs, annual or biennial, to 1.2(–2.4) m. Leaves mesophyllous, 3.5–17 cm, pallid green, slightly and finely hairy or glabrous; stipules caducous; extrafloral nectary 1, usually between first leaflet pair, rarely also between second, sessile or shortly stipitate; leaflet pairs 3, blades obovate to cuneate-obovate or broadly cuneate-oblanceolate, 17–65 × 10–40 mm.
Racemes

1–5-flowered;

bracts caducous.

usually 1 or 2(or 3)-flowered;

bracts caducous.

Pedicels

9–16 mm.

7–28 mm.

Flowers

monosymmetric;

calyx caducous, pale green;

corolla yellow or orange-yellow, longest petal 12–17 mm;

androecium not heterantherous, stamens 7, staminodes 3;

anthers 2.2–3.3 mm, dehiscing by 1 apical pore, apical appendage 0;

gynoecium nearly linear, ovules 22–40;

ovary densely hairy;

style slightly incurved.

asymmetric, enantiostylous;

calyx pale green;

corolla pale yellow, longest petal 9–15 mm, 1 lower petal conspicuously larger;

androecium heterantherous, stamens 7, middle stamens 1/2 as long as abaxial or smaller, staminodes 3;

anthers of middle stamens 1–2.8 mm, of abaxial stamens 2–5 mm, dehiscing by U-shaped slit, apical appendage inconspicuous, cupped;

gynoecium incurved, ovules 16–38;

ovary hairy;

style incurved.

Legumes

erect, subcylindrical, straight or slightly curved, 20–35 × 4.5–6.5 mm, corrugated over seeds, dehiscing apically downward.

erect or curved downward, flat, straight, 60–180 × 2.5–6 mm, faintly corrugated over seeds, indehiscent.

Seeds

brown or pinkish brown, paddle-shaped or pyriform.

dark reddish brown, rhomboid or subcylindroid-oblong.

2n

= 28.

= 24, 26, 28.

Senna roemeriana

Senna obtusifolia

Phenology Flowering spring–early fall. Flowering late spring–mid winter.
Habitat Mesquite grasslands, chaparral, draws in shortgrass prairies, barren hillsides, desert washes, roadsides. Lakeshores, riverbanks, river beds, disturbed habitats, pastures, plantations, orchards, roadsides, waste places.
Elevation 100–2000 m. (300–6600 ft.) 0–1700 m. (0–5600 ft.)
Distribution
from FNA
NM; OK; TX; Mexico (Chihuahua, Coahuila, Nuevo León)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; FL; GA; IL; IN; KS; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OK; PA; SC; TN; TX; VA; WV [Introduced in Asia, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Senna obtusifolia is one of the most widespread weedy sennas native to the Americas (H. S. Irwin and R. C. Barneby 1982) and is probably naturalized circumtropically. The species is considered a noxious weed in many countries, posing problems especially for agriculture; it is able to completely invade pastures by dominating grass species, it strongly competes with crops, affecting yields negatively, and, although generally unpalatable to stock, if eaten, it is toxic to cattle. For these reasons, in Australia S. obtusifolia is designated as potentially of national concern, and authorities estimate that this invader can lead properties to become unproductive (Weeds of Australia, www.environment.gov.au/cgi-bin/biodiversity/invasive/weeds/weedsearch.pl; Land Protection, www.nrw.qld.gov.au/factsheets/pdf/pest/pp18.pdf). In the flora area, its range of distribution appears to have expanded from five southern and southeastern states in the early 1980s (Florida, Kentucky, Missouri, Texas, and Virginia; Irwin and Barneby) to currently almost half of the states. Although a few specimens were collected as far north as Nebraska and Wisconsin, it is not known to be established in those states.

Senna tora (Linnaeus) Roxburgh (Cassia tora Linnaeus) is sometimes considered synonymous with S. obtusifolia. However, due to differences in length and curvature of the fruit and in length of the petiole and pedicel, B. R. Randell and B. A. Barlow (1998) considered it distinct. H. S. Irwin and R. C. Barneby (1982) observed a range rather than distinct categories in the measures and shapes of these traits, and only molecular phylogenetic and biogeographic studies accompanied by morphometric analyses may definitively solve this taxonomic dilemma.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Caesalpinioideae (excluding Mimosoid clade) > Senna Fabaceae > subfam. Caesalpinioideae (excluding Mimosoid clade) > Senna
Sibling taxa
S. alata, S. armata, S. artemisioides, S. atomaria, S. bauhinioides, S. corymbosa, S. covesii, S. durangensis, S. hebecarpa, S. hirsuta, S. ligustrina, S. lindheimeriana, S. marilandica, S. mexicana, S. multiglandulosa, S. obtusifolia, S. occidentalis, S. orcuttii, S. pendula, S. pilosior, S. pumilio, S. ripleyana, S. surattensis, S. wislizeni
S. alata, S. armata, S. artemisioides, S. atomaria, S. bauhinioides, S. corymbosa, S. covesii, S. durangensis, S. hebecarpa, S. hirsuta, S. ligustrina, S. lindheimeriana, S. marilandica, S. mexicana, S. multiglandulosa, S. occidentalis, S. orcuttii, S. pendula, S. pilosior, S. pumilio, S. ripleyana, S. roemeriana, S. surattensis, S. wislizeni
Synonyms Cassia roemeriana, Earleocassia roemeriana Cassia obtusifolia, C. tora var. humilis, C. tora var. obtusifolia, C. toroides
Name authority (Scheele) H. S. Irwin & Barneby: Mem. New York Bot. Gard. 35: 282. (1982) (Linnaeus) H. S. Irwin & Barneby: Mem. New York Bot. Gard. 35: 252. (1982)
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