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flor de San Jose, flor de san josé, palo zorillo

blunt leafed senna, Chinese senna, Java-bean, sickle-pod wild senna, sicklepod

Habit Shrubs or trees, to 20 m. Leaves mesophyllous to slightly sclerophyllous, 8.5–28.5 cm, hairy, sometimes densely; stipules caducous; extrafloral nectaries 0; leaflet pairs 2–5, blades bicolored, usually obovate to elliptic, sometimes ovate, 20–130 × 10–60 mm. Herbs, annual or biennial, to 1.2(–2.4) m. Leaves mesophyllous, 3.5–17 cm, pallid green, slightly and finely hairy or glabrous; stipules caducous; extrafloral nectary 1, usually between first leaflet pair, rarely also between second, sessile or shortly stipitate; leaflet pairs 3, blades obovate to cuneate-obovate or broadly cuneate-oblanceolate, 17–65 × 10–40 mm.
Racemes

5–55-flowered, not spikelike;

bracts early caducous, to 5 mm.

usually 1 or 2(or 3)-flowered;

bracts caducous.

Pedicels

13–28 mm.

7–28 mm.

Flowers

asymmetric, enantiostylous;

calyx greenish to yellow;

corolla yellow-orange, slightly dark-veined, longest petal 12–23 mm, highly asymmetric, 1 or both lower petals highly modified, strongly concave and folded over stamens (flag-shaped);

androecium slightly heterantherous, stamens 7 (similar in shape and size, abaxial ones slightly longer), staminodes 3;

anthers 2.8–5 mm, dehiscing by 2 short slits, apical appendage 0;

gynoecium incurved, ovules 46–70;

ovary glabrate, sometimes becoming hairy after fertilization;

style stout.

asymmetric, enantiostylous;

calyx pale green;

corolla pale yellow, longest petal 9–15 mm, 1 lower petal conspicuously larger;

androecium heterantherous, stamens 7, middle stamens 1/2 as long as abaxial or smaller, staminodes 3;

anthers of middle stamens 1–2.8 mm, of abaxial stamens 2–5 mm, dehiscing by U-shaped slit, apical appendage inconspicuous, cupped;

gynoecium incurved, ovules 16–38;

ovary hairy;

style incurved.

Legumes

pendulous, flat, straight, 220–370 × 80–140 mm, woody, indehiscent or splitting transversely into woody segments.

erect or curved downward, flat, straight, 60–180 × 2.5–6 mm, faintly corrugated over seeds, indehiscent.

Seeds

reddish brown, obovoid to oblong-obovoid.

dark reddish brown, rhomboid or subcylindroid-oblong.

2n

= 24, 26, 28.

Senna atomaria

Senna obtusifolia

Phenology Flowering late winter–late spring. Flowering late spring–mid winter.
Habitat Disturbed habitats. Lakeshores, riverbanks, river beds, disturbed habitats, pastures, plantations, orchards, roadsides, waste places.
Elevation 0–20 m. (0–100 ft.) 0–1700 m. (0–5600 ft.)
Distribution
from FNA
FL; Mexico (Baja California Sur, Campeche, Chiapas, Colima, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, Quintana Roo, San Luis Potosí, Sinaloa, Sonora, Tabasco, Tamaulipas, Veracruz, Yucatán); Central America (including Caribbean Islands); South America (Colombia, Ecuador, Venezuela) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; FL; GA; IL; IN; KS; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OK; PA; SC; TN; TX; VA; WV [Introduced in Asia, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

As with other trees from deciduous and semi-deciduous vegetation, Senna atomaria is covered with flowers before developing the foliage (H. S. Irwin and R. C. Barneby 1982). In the flora area, it occurs naturalized only very locally in Collier County (R. P. Wunderlin et al., http://florida.plantatlas.usf.edu/).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Senna obtusifolia is one of the most widespread weedy sennas native to the Americas (H. S. Irwin and R. C. Barneby 1982) and is probably naturalized circumtropically. The species is considered a noxious weed in many countries, posing problems especially for agriculture; it is able to completely invade pastures by dominating grass species, it strongly competes with crops, affecting yields negatively, and, although generally unpalatable to stock, if eaten, it is toxic to cattle. For these reasons, in Australia S. obtusifolia is designated as potentially of national concern, and authorities estimate that this invader can lead properties to become unproductive (Weeds of Australia, www.environment.gov.au/cgi-bin/biodiversity/invasive/weeds/weedsearch.pl; Land Protection, www.nrw.qld.gov.au/factsheets/pdf/pest/pp18.pdf). In the flora area, its range of distribution appears to have expanded from five southern and southeastern states in the early 1980s (Florida, Kentucky, Missouri, Texas, and Virginia; Irwin and Barneby) to currently almost half of the states. Although a few specimens were collected as far north as Nebraska and Wisconsin, it is not known to be established in those states.

Senna tora (Linnaeus) Roxburgh (Cassia tora Linnaeus) is sometimes considered synonymous with S. obtusifolia. However, due to differences in length and curvature of the fruit and in length of the petiole and pedicel, B. R. Randell and B. A. Barlow (1998) considered it distinct. H. S. Irwin and R. C. Barneby (1982) observed a range rather than distinct categories in the measures and shapes of these traits, and only molecular phylogenetic and biogeographic studies accompanied by morphometric analyses may definitively solve this taxonomic dilemma.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Caesalpinioideae (excluding Mimosoid clade) > Senna Fabaceae > subfam. Caesalpinioideae (excluding Mimosoid clade) > Senna
Sibling taxa
S. alata, S. armata, S. artemisioides, S. bauhinioides, S. corymbosa, S. covesii, S. durangensis, S. hebecarpa, S. hirsuta, S. ligustrina, S. lindheimeriana, S. marilandica, S. mexicana, S. multiglandulosa, S. obtusifolia, S. occidentalis, S. orcuttii, S. pendula, S. pilosior, S. pumilio, S. ripleyana, S. roemeriana, S. surattensis, S. wislizeni
S. alata, S. armata, S. artemisioides, S. atomaria, S. bauhinioides, S. corymbosa, S. covesii, S. durangensis, S. hebecarpa, S. hirsuta, S. ligustrina, S. lindheimeriana, S. marilandica, S. mexicana, S. multiglandulosa, S. occidentalis, S. orcuttii, S. pendula, S. pilosior, S. pumilio, S. ripleyana, S. roemeriana, S. surattensis, S. wislizeni
Synonyms Cassia atomaria, C. emarginata Cassia obtusifolia, C. tora var. humilis, C. tora var. obtusifolia, C. toroides
Name authority (Linnaeus) H. S. Irwin & Barneby: Mem. New York Bot. Gard. 35: 588. (1982) (Linnaeus) H. S. Irwin & Barneby: Mem. New York Bot. Gard. 35: 252. (1982)
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