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silver senna

trans-Pecos senna

Habit Shrubs, to 3 m. Leaves slightly to highly xerophytic as phyllodes, 0.8–5 cm, finely hairy; stipules caducous; extrafloral nectaries 1–8, between first leaflet pair, sometimes also 1 or all subsequent pairs, sessile; leaflet pairs 0–8, blades linear to narrowly elliptic, 20–70 × 1–8 mm. Herbs, perennial, to 0.7 m. Leaves slightly sclerophyllous, 1.5–7.5 cm, hairy; stipules tardily deciduous; extrafloral nectary 1, between leaflet pair, stipitate; leaflet pairs 1, blades broadly, obliquely obovate to oblong-obovate, 15–40 × 10–25 mm.
Racemes

2–10-flowered;

bracts early caducous.

2–6-flowered;

bracts caducous.

Pedicels

4–15 mm.

6–15 mm.

Flowers

slightly asymmetric, enantiostylous;

calyx yellowish to greenish;

corolla yellow, longest petal 6–10 mm;

androecium not heterantherous, stamens 10, staminodes 0;

anthers 1.5–5 mm, dehiscing by 2 pores, apical appendage 0;

gynoecium incurved, ovules unknown;

ovary slightly hairy;

style slightly incurved.

monosymmetric;

calyx persistent into developing fruit, pale green;

corolla yellow, longest petal 8.5–10 mm;

androecium not heterantherous, stamens 7, staminodes 3;

anthers 2.3–3.7 mm, dehiscing by 1 apical pore, apical appendage 0;

gynoecium nearly linear, ovules 16–26;

ovary densely hairy;

style linear.

Legumes

pendulous, flat, straight or slightly curved, 20–80 × 6–12 mm, corrugated over seeds, indehiscent.

erect, flat, straight, 20–40 × 5.5–7.5 mm, corrugated over seeds, dehiscing apically downward, with 1 series of seeds.

Seeds

dark brown or dull, obovoid.

dull pinkish or grayish brown, pyriform.

2n

= 28, 56.

= 28.

Senna artemisioides

Senna pilosior

Phenology Flowering late fall–early spring. Flowering spring–mid fall.
Habitat Rocky and desert sand and soils. Sandy banks, desert washes.
Elevation 0–700 m. (0–2300 ft.) 600–1500 m. (2000–4900 ft.)
Distribution
from FNA
AZ; CA; Australia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Chihuahua, Coahuila, Durango)
[BONAP county map]
Discussion

Due to its remarkable morphological and genetic variation (especially in leaflet number and form), Senna artemisioides has a shifting taxonomic history. The species, as circumscribed by Randell, was divided into several morphological forms by B. R. Randell and B. A. Barlow (1998). Senna artemisioides, as presented here, encompasses four common morphological forms found in North America: subsp. filifolia Randell, subsp. petiolaris Randell, nothosubsp. sturtii (R. Brown) Randell, and subsp. zygophylla Randell. D. E. Symon (1998) considered individuals of subsp. petiolaris characterized by arcuate phyllodes to be a separate species, S. phyllodinea (R. Brown) Symon, mainly because of the absence of individuals with an intermediate leaf morphology transitioning to the phyllode shape. Although this argument is reasonable, S. phyllodinea has not been accepted and adopted in other floristic studies. It must be noted that phyllodes consist of the lateral compression of both the petiole and the rachis, and not only of the petiole, as reported by Randell and Barlow. Along the adaxial margin of the phyllode, there is, in fact, a reduced extrafloral nectary at each insertion place of the suppressed leaflets.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Senna pilosior is the only senna in North America with sepals persisting into fruit development and falling off before fruit matures and dehisces. Persistent sepals are diagnostic to distinguish S. pilosior from the close relatives S. bauhinioides and S. durangensis; the trio is, in fact, characterized by a history of repeated misidentifications (H. S. Irwin and R. C. Barneby 1982).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Caesalpinioideae (excluding Mimosoid clade) > Senna Fabaceae > subfam. Caesalpinioideae (excluding Mimosoid clade) > Senna
Sibling taxa
S. alata, S. armata, S. atomaria, S. bauhinioides, S. corymbosa, S. covesii, S. durangensis, S. hebecarpa, S. hirsuta, S. ligustrina, S. lindheimeriana, S. marilandica, S. mexicana, S. multiglandulosa, S. obtusifolia, S. occidentalis, S. orcuttii, S. pendula, S. pilosior, S. pumilio, S. ripleyana, S. roemeriana, S. surattensis, S. wislizeni
S. alata, S. armata, S. artemisioides, S. atomaria, S. bauhinioides, S. corymbosa, S. covesii, S. durangensis, S. hebecarpa, S. hirsuta, S. ligustrina, S. lindheimeriana, S. marilandica, S. mexicana, S. multiglandulosa, S. obtusifolia, S. occidentalis, S. orcuttii, S. pendula, S. pumilio, S. ripleyana, S. roemeriana, S. surattensis, S. wislizeni
Synonyms Cassia artemisioides Cassia bauhinioides var. pilosior, C. pilosior
Name authority (Gaudichaud-Beaupré ex de Candolle) Randell: J. Adelaide Bot. Gard. 12: 220. (1989) (B. L. Robinson ex J. F. Macbride) H. S. Irwin & Barneby: Phytologia 44: 500. (1979)
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