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selenia

Jones' selenia

Habit Annuals (sometimes winter); not scapose; glabrous. Plants winter annuals, (not or rarely subacaulescent).
Stems

(rarely absent, base of plant forming inflated crown), erect, ascending, subdecumbent, or decumbent, unbranched or branched distally.

(not inflated into crown), ascending or subdecumbent, (0.5–)1–3(–4) dm.

Leaves

basal and cauline;

petiolate or subsessile;

basal rosulate or not, petiolate, blade margins 1- or 2- (or 3-)pinnatisect, (terminal lobe margin entire or dentate);

cauline sessile [subsessile], blade (base not auriculate), margins similar to basal.

Basal leaves

rosulate;

petiole 1–2.5 cm;

blade margins 2-pinnatisect, 4–8 cm;lobes 4–11 on each side, (smaller than terminal);

apical segment linear to oblong or ovate, 1–6 × 0.5–2.5 mm, margins entire.

Cauline leaves

(and bracts) similar to basal, smaller distally.

Racemes

(sometimes pedicels originating between basal leaves, bracteate throughout, rachis straight), considerably elongated in fruit.

Flowers

sepals (sometimes persistent), erect, spreading, or ascending, (yellowish), oblong, oblong-linear or -lanceolate, lateral pair slightly saccate basally;

petals yellow, spatulate to broadly obovate, claw differentiated from blade, (apex rounded or emarginate [obtuse]);

stamens tetradynamous;

filaments not dilated basally;

anthers linear, ovate, or oblong [sagittate];

nectar glands confluent (extending into spreading teeth).

sepals spreading, oblong, 4–6(–7) × 1.5–2.5 mm, apex appendage developed, 0.5–0.8 mm;

petals spatulate, 4–7 × 2.5–4 mm, apex rounded or emarginate;

median filament pairs 3–4 mm, not dilated basally;

anthers ovate, 1–1.5 mm;

gynophore 1–3 mm.

Fruiting pedicels

divaricate to ascending, slender.

some from basal leaf axils, (10–)20–40(–50) mm.

Fruits

silicles, sessile or shortly stipitate, usually oblong, elliptical, suborbicular, or globose, rarely obovoid, smooth, latiseptate, terete, or slightly inflated;

valves (vesicular in S. grandis, thin-papery to leathery), each without midvein, obscurely to prominently reticulate-veined;

replum rounded or flattened;

septum complete or perforated;

ovules 8–44 per ovary;

style distinct, (1–12 mm, sometimes flattened basally);

stigma capitate.

globose or, rarely, obovoid, terete, 0.8–1.5 cm × 8–14 mm, (not fleshy, papery), base usually obtuse, rarely cuneate, apex obtuse;

valves obscurely reticulate-veined;

replum not flattened;

septum complete;

ovules 8–14 per ovary;

style 1–2(–3) mm, not flattened basally.

Seeds

biseriate, flattened, winged throughout, orbicular;

seed coat (coarsely reticulate), not mucilaginous when wetted;

cotyledons accumbent.

4–5 mm diam.;

wing 1–1.5 mm.

x

= 7, 12, 13.

2n

= 24.

Selenia

Selenia jonesii

Phenology Flowering Mar–Apr.
Habitat Dry lake beds, draws, moist swales, prairie plateaus, playa lakes, buffalo wallows
Elevation 0-1100 m (0-3600 ft)
Distribution
from USDA
c United States; sw United States; ne Mexico
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Species 5 (4 in the flora).

Species of Selenia might be difficult to separate when the plants are only in flower. Selenia aurea is easily distinguished by its 1-pinnatisect leaves and absence of flowers from the base of the plant. The remaining three species have 2- or 3-pinnatisect leaves and at least some pedicels from the basal rosette. In S. jonesii, the sepal appendage is absent or less than 1 mm; in both S. dissecta and S. grandis, it is 1–4 mm. The last two can be distinguished in flower by the presence in S. grandis of terete styles and ovary margins (replum) and in S. dissecta by flattened style bases and winged ovary margins. Selenia mexicana Standley is endemic to Nuevo León, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Selenia jonesii is endemic to six counties in the western Edwards Plateau. Variety obovata is reduced herein to synonymy because the alleged difference in fruit shape does not hold; it appears to be an artifact of pressing inflated fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade margins 1-pinnatisect; fruiting pedicels from racemes; sepals with unappendaged apex.
S. aurea
1. Leaf blade margins 2- or 3-pinnatisect; fruiting pedicels (at least some) from basal leaf axils or rosettes; sepals with appendaged apex
→ 2
2. Fruit valves vesicular, leathery; sepals persistent; fruits 5.5-7.5 mm wide.
S. grandis
2. Fruit valves not vesicular, papery; sepals caducous or tardily so; fruits 8-17 mm wide
→ 3
3. Fruits oblong to elliptical, latiseptate; replums and styles flattened; petals (12-)15-20 × (5-)6-9 mm; ovules 28-40 per ovary; median filament pairs 6-10 mm; sepal apex appendage (1-)1.5-3 mm.
S. dissecta
3. Fruits usually globose, rarely obovoid, terete; replums and styles not flattened; petals 4-7 × 2.5-4 mm; ovules 8-14 per ovary; median filament pairs 3-4 mm; sepal apex appendage 0.5-0.8 mm.
S. jonesii
Source FNA vol. 7, p. 506. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 508.
Parent taxa Brassicaceae > tribe Cardamineae Brassicaceae > tribe Cardamineae > Selenia
Sibling taxa
S. aurea, S. dissecta, S. grandis
Subordinate taxa
S. aurea, S. dissecta, S. grandis, S. jonesii
Synonyms S. jonesii var. obovata
Name authority Nuttall: J. Acad. Nat. Sci. Philadelphia 5: 132, plate 6. (1825) Cory: Rhodora 33: 142. (1931)
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