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Underwood's spike-moss

green spikemoss, slender spike-moss

Habit Plants on rock, forming loose festoonlike mats or rarely compact mats. Plants on rock, forming clumps or mounds.
Stems

radially symmetric, long-creeping, short-creeping, or pendent, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight;

main stem indeterminate, lateral branches determinate, spreading, 1–2-forked.

radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked; rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight;

aerial stems mainly erect, seldom ascending, with budlike arrested branches throughout stem length.

Leaves

monomorphic, in alternate pseudowhorls of 4 (on main stem and older lateral branches) or 3 (on young lateral branches and secondary branches), loosely appressed, ascending, green, linear to linear-lanceolate or narrowly triangular-lanceolate, (2–)2.5–3.4 × 0.45–0.5(–0.7) mm;

abaxial ridges prominent;

base mostly cuneate and decurrent, rarely rounded and adnate (on young branches), pubescent or glabrous;

margins entire to denticulate or very short-ciliate, cilia transparent, scattered, mostly ascending, dentiform toward apex, 0.02–0.07 mm;

apex keeled, slightly attenuate, short- to long-bristled;

bristle transparent greenish to greenish-yellowish, rarely white, smooth, seldom slightly puberulent, sometimes breaking off, 0.25–0.7(–1) mm.

dimorphic, not clearly ranked.

Strobili

sometimes paired, 0.5–3.5 cm;

sporophylls lanceolate to ovate-lanceolate, abaxial ridges prominent, base glabrous, with prominent auricles (no other species has such prominent auricles), margins entire or very short-ciliate to denticulate, apex keeled, short- to long-bristled.

solitary, 0.5–1.2(–2.5) cm;

sporophylls deltate-ovate to ovate-lanceolate, abaxial ridges prominent, base glabrous, margins denticulate, apex acute to obtuse.

Rhizophores

borne on upperside of stems, throughout stem length, 0.15–0.27(–0.3) mm diam.

borne on upperside of stems, restricted to rhizomatous stems and lower 1/4 of aerial stems, 0.16–0.3 mm diam.

Rhizomatous

stem leaves loosely appressed, straight, scalelike.

Aerial

stem leaves appressed, ascending, green, linear-lanceolate to narrowly lanceolate, 1.8–2.1 × 0.49–0.56 mm;

abaxial ridges prominent;

base cuneate and decurrent to slightly rounded and adnate, glabrous;

margins denticulate to very short-ciliate, cilia transparent, spreading to ascending toward apex, 0.02–0.04 mm;

apex acute or seldom blunt.

Selaginella underwoodii

Selaginella viridissima

Habitat Moist or shaded cliffs, rocky slopes, rock crevices, granitic outcrops, hanging over granite cliffs, sandstone or limestone ledges Shaded cliffs, slopes, rock crevices, and igneous rock
Elevation (800–)1500–3000(–4000) m ((2600–)4900–9800(–13100) ft) 1650–2300 m (5400–7500 ft)
Distribution
from FNA
AZ; CO; NM; OK; TX; UT; WY; Mexico in Chihuahua and Nuevo León
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico in Coahuila
[BONAP county map]
Discussion

R. M. Tryon (1971) reported that the bristle on Selaginella underwoodii leaves is longer (to 1.44 mm) in the southern part of the range and shorter (to 0.43 mm) northward and in central Arizona. Selaginella underwoodii seems to be closely related to S. oregana, perhaps sharing a common ancestor.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Texas Selaginella viridissima is known only from the Chisos Mountains.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Selaginellaceae > Selaginella > subg. Tetragonostachys Selaginellaceae > Selaginella > subg. Tetragonostachys
Sibling taxa
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
Synonyms S. fendleri S. coryi
Name authority Hieronymus: in Engler & Prantl, Nat. Pflanzenfam. 1: 714. (1901) Weatherby: J. Arnold Arbor. 24: 326. (1943)
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