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rockloving spikemoss

ashy spike-moss, gray spike-moss, mesa spikemoss

Habit Plants on rock or terrestrial, forming loose clumps. Plants terrestrial, forming loose to compact mats.
Stems

radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked;

both rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight; rhizomatous stems hard to distinguish on wholly creeping plants;

aerial stems erect or ascending, sometimes decumbent to slightly creeping, budlike arrested branches restricted mostly near stem base.

creeping, not readily fragmenting, upperside and underside structurally slightly different, irregularly forked, without budlike arrested branches, tips straight;

main stem indeterminate, lateral branches determinate, ascending, 1–2-forked.

Leaves

dimorphic, not clearly ranked.

monomorphic, not in defined pseudowhorls, loosely appressed, ascending, green, linear-lanceolate, (1–)2.5–3 X (0.25–)0.4–0.6 mm (leaves in secondary and tertiary branches smaller);

abaxial ridges inconspicuous;

base rounded and adnate or cuneate and slightly decurrent, glabrous, seldom pubescent;

margins short-ciliate, cilia transparent, scattered, ascending, 0.02–0.75 mm;

apex plane, blunt, acute to slightly acuminate (not distinctly bristled).

Strobili

solitary, 0.5–2.5(–3.5) cm;

sporophylls lanceolate, strongly tapering toward tip, abaxial ridges prominent, base glabrous, margins short-ciliate, apex long-bristled.

solitary, 2–4 mm;

sporophylls deltate-ovate to lanceolate-ovate, abaxial ridges not prominent, base glabrous, margins short-ciliate, apex not keeled, acute.

Rhizophores

borne on upperside of stems, restricted to lower stems or throughout stem length, 0.3–0.5 mm diam.

borne on upperside of stems, throughout stem length, (0.17–)0.2–0.3 mm diam.

Rhizomatous

stem leaves persistent or deciduous, tightly appressed, scalelike.

Aerial

stem leaves appressed, ascending, green, linear-lanceolate, 3–4.7 × 0.45–0.65 mm;

abaxial ridges present;

base abruptly adnate, rounded, pubescent;

margins long-ciliate, cilia white to whitish, spreading, 0.1–0.2 mm;

apex not keeled to slightly keeled;

bristle white to whitish or yellowish to greenish near base, puberulent, 0.65–1.85 mm (1/3–1/2 length of leaves).

Selaginella rupincola

Selaginella cinerascens

Habitat Exposed ledges and rock, steep slopes, rock crevices or gravelly soil Dry open places of clay soil, clayey-sandy soil, or in shade under shrubs and trees
Elevation 1000–2000 m (3300–6600 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AZ; NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

Selaginella rupincola is allied to S. bigelovii. It is one of the presumed parents of S. × neomexicana (see discussion). In addition to characteristics given, it can be separated from S. bigelovii in having hairs often running along the ridges of the abaxial groove, whereas S. bigelovii has nonhairy ridges on the abaxial groove.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The light brown and grayish mats, short lateral branches, narrow stem, and short strobili distinguish Selaginella cinerascens from all other species in the flora, in which it has no close relatives. R. M. Tryon (1955) related S. cinerascens to S. arsenei Weatherby from Mexico. Selaginella cinerascens also closely resembles S. nivea Alston from Madagascar. In California S. cinerascens is known only from San Diego County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Selaginellaceae > Selaginella > subg. Tetragonostachys Selaginellaceae > Selaginella > subg. Tetragonostachys
Sibling taxa
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. cinerascens, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
S. acanthonota, S. apoda, S. arenicola, S. arizonica, S. asprella, S. bigelovii, S. braunii, S. densa, S. douglasii, S. eatonii, S. eclipes, S. eremophila, S. hansenii, S. kraussiana, S. lepidophylla, S. leucobryoides, S. ludoviciana, S. mutica, S. oregana, S. peruviana, S. pilifera, S. rupestris, S. rupincola, S. scopulorum, S. selaginoides, S. sibirica, S. standleyi, S. tortipila, S. uncinata, S. underwoodii, S. utahensis, S. viridissima, S. wallacei, S. watsonii, S. weatherbiana, S. willdenowii, S. wrightii, S. ×neomexicana
Name authority L. Underwood: Bull. Torrey Bot. Club 25: 129. (1898) A. A. Eaton: Fern Bull. 7: 33. (1899)
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